The Inheritance of Peloria in Foxgloves. 69 
1. Peloria in Foxgloves. Turning now to our experiments 
with Foxgloves: all interested in gardens are aware that races of 
these plants are in cultivation which exhibit the phenomenon of 
peloria. In Foxgloves, peloria, at all events in the examples which 
we have had under observation, is confined to the terminal flower 
of the raceme. Whereas the lateral flowers retain their typical 
zygomorphic state, the terminal flower of a peloric plant is bell¬ 
shaped. Moreover, it stands up vertically and so forms a striking 
contrast with the pendulous, zygomorphic, lateral flowers. The 
peloric flower shows other indications of its monstrous nature ; its 
corolla may he nine-lobed, the anthers as many as 8, and the 
stigma possessed of 5 stigmatic surfaces. 
Though built on such abnormal and massive lines the peloric 
flowers of Foxgloves appear to he perfectly fertile with their own 
pollen. Thus the terminal, regular flowers of a peloric plant (plant B, 
Table I, p. 76) self-pollinated in 1907, gave seed from which 12 
plants were raised. These 12 flowered in 1909 and were all peloric. 
In the next place it is noteworthy that the typical, zygomorphic, 
lateral flowers of peloric plants carry, though they do not reveal, the 
factor which determines peloria. For example; the lateral 
flowers of a peloric plant (Plant A) selfed, gave an F, of 22 plants 
all peloric with respect to their terminal flowers. The question as to 
the significance of the facts just mentioned is one on which we 
are at present engaged ; but with respect to which we are not at 
present prepared to make any statement. That peloria depends 
upon a recessive character is indicated by the following records:— 
Plant D : non-peloric, selfed, 1907. 
F,=ll non-peloric and 3 peloric. 
Plant E : non-peloric, selfed, 1907. 
F,=9 non-peloric and 4 peloric. 
Combining the numbers obtained in these experiments we have 
non-peloric, 20, peloric, 7. From which we may conclude that peloria 
(p) is recessive to normal (P). Crosses between peloric and non- 
peloric confirm this conclusion. Thus plant D (non-peloric) (Pp) 
X plant E (non-peloric) (Pp) F t should give:— 
1 PP : 2 Pp : 1 pp. 
3 Dominant : 1 Recessive. 
Our numbers=35 Dominant : 11 Recessive, 
