70 F. Keeble, Miss C. Pellew and W. N. Jones. 
Another case: plant F (peloric) (pp) X plant E (non peloric) 
(Pp) should give :— 
2 pp : 2 Pp. 
Observed :— 16 : 24. 
Peloria of Foxgloves therefore depends on a character which 
is recessive to normal. Inasmuch as it is characterised primarily 
by absence of the typical zygomorphy of normal flowers, and 
inasmuch as evidence is accumulating in support of the presence 
and absence hypothesis of the nature of dominant and recessive 
members of allelomorphic pairs of characters, it is not perhaps 
surprising to find peloria behaving as recessive to normal. 
2. Flower Colour. The colour-characteristics, the mode of 
inheritance of which we are investigating, are white, magenta, deep 
magenta : red spots, brown and yellow spots, which characters 
are displayed by the corollas of various garden races of Foxgloves. 
In Tables I and II (pp. 76, 77) are tabulated the colour-characters 
of the various plants under investigation and the behaviour of these 
characters in the F t generations resulting from self-fertilization of 
these plants, and from the cross-fertilization of plant D x plant E 
and plant F x plant E. 
Inspection of Table I. shows that certain white plants breed 
true to whiteness (Plant E) ; others, e.g., plant D, when selfed, 
yield an F, composed of w r hite and magenta-flowered plants. 
Similarly when E, which, so far as the comparatively small numbers 
in F x admit of a conclusion, is to be regarded as pure to whiteness, 
is crossed with D, whites and magentas are obtained. We therefore 
conclude that Foxgloves, like Primula sinensis (Bateson, 1909), 
contain a dominant-white factor (W), that is a factor which inhibits 
the expression of colour in a zygote, the contributory gametes of 
which carried a colour factor. Since colour appears in F, from 
plant D selfed, that plant is not pure, but heterozygous with respect 
to the dominant white factor, i.e., its gametic constitution is Ww. 
Nor may the plant D be pure with respect to magenta colour-factor 
(M), for, if it were, its gametic constitution would be Ww MM ; it 
would yield gametes wM and WM and, on selfing, would produce 
an F, composed of 3 dominant white : 1 magenta, thus :— 
WM.wM, male gametes. 
WM.wM, female ,, 
giving in Fj 
1 WW MM : 2 Ww MM : 1 ww MM 
> -- - ^ 
3 white : 1 magenta. 
