75 
The Inheritance of Peloria in Foxgloves. 
Plant F (Ww MM) selfed, gives P p all with red spots. 
Plant E (ww min) selfed, gives F x , all with yellow-brown spots . 
Plant F x E (Ww MM X ww mm) gives F lf all with red 
spots (40 plants). 
Plant D (Ww Mm) selfed, gives Fj, 12 plants with red, 2 with 
brown spots. 
Plant A (ww Mm) selfed, gives F 1? 17 plants with red, 5 with 
brown spots. 
Totals—29 plants with red, 7 with brown spots. 
Calculated—27 plants with red, 9 with brown spots. 
It should be noted that in the results of plant D selfed there is 
a distinct departure from the expected numbers, viz :— 
Obtained=12 : 2 
Calculated=10-5 : 3-5 
That this is, however, attributable to the fewness of the Fj 
plants is rendered probable by the results obtained by crossing 
plants D and E. 
Plants D x E=Ww Mm x ww mm should yield F, composed 
of 4 Mm ww : 4 Mm Ww : 4 mm Ww : 4 mm ww, that is equal 
numbers of red spotted and yellow-brown spotted flowers. The 
numbers obtained are 21 plants with red spotted, 25 with brown 
spotted flowers. 
The hypothesis that red spots are due to magenta factor acting 
on yellow-brown spot-areas provides a simple explanation of the 
relation between colour of spots and ground colour exhibited by the 
F x from plant A ; viz., 17 magenta-containing plants all with red 
spots and 6 white plants all with yellow-brown spots. It also 
explains how it is that all the magenta Foxgloves have red spots. 
It is not improbable that the view which we have expressed, namely 
that a dominant white factor (W) is able to inhibit ground colour 
and not spot colour—though both ground colour and spot colour are 
due to the same colour factor (M),—may prove of importance in 
interpreting the origin of spots and bars and stripes in animals and 
plants. Given localised inconspicuous areas, the addition of a ground 
colour factor would produce a self (uniformly) coloured surface on 
which spots and bars and stripes might or might not stand out 
conspicuously. Wash over that surface with a dominant white 
factor and then the general ground colour will be suppressed and 
conspicuous splashes of colour in spots, colours or stripes will stand 
out on a white ground. Finally, the suggestion may be worth making, 
that this differential action of a dominant white factor—inhibiting 
