258 Two Embryo-Sacs in Fritillaria. 
cells were found. Other cases mentioned in this note are Liliutn 
philadelphicum with three and five archesporial cells found by 
Coulter and Chamberlain 1 and that mentioned by Bernard 2 in Liliutn 
candidum where two embryo-sacs, showing the stage with four 
nuclei, were found lying side by side in intimate contact along their 
inner faces. He thinks from their appearance they may have arisen 
by division of a single archesporial cell. 
The case described by Miss Ferguson, however, shows two 
distinct archesporial cells separated by a layer of tissue of the 
nucellus. They are surrounded by the usual single layer of outer 
cells of the nucellus, and normal integuments are present. The 
nuclei as figured show the chromatin thread and nucleoli in resting 
condition before commencement of the meiotic division. 
Fig. 19. 
The case now to be described shows an intermediate condition 
between the cases described by Bernard and Miss Ferguson. 
In this case the two embryo-sac mother-cells are seen lying 
side by side in close contact along their inner faces, but with a 
distinct wall between them. 
The two embryo-sacs seem from their appearance to have 
originated from two distinct archesporial cells. They are enclosed 
by the usual single layer of cells of the nucellus, and the ovule has 
normal integuments. The nuclei show the stage following synapsis, 
the spireme thread has broken up into chromosomes previous to 
the arrangement on the equatorial plane. The section is, unfortu¬ 
nately, rather oblique and only one nucleus can be seen in detail, 
but it must be inferred that each nucleus will show the same 
condition, and as the stage shown by the one nucleus is that pre¬ 
vious to the reduction division it is obvious that each embryo sac 
must have arisen from a separate archesporial cell, as in Fritil¬ 
laria the first division of the nucleus of the archesporial cell is 
the reduction division. 
1 Coulter and Chamberlain. “ Morphology of Angiospcrms.” 
New York, 1903, p. 61. 
3 Bernard, C. H. “ Recherches sur les spheres attractives chez 
Liliutn candidum." 1900. Jour, de Botanique 14, p. 178. 
