320 
F. Summers. 
THE LEAF OF COLLIQUAYA ODOHIFERA , MOLIN. 
By F. Summers, B.Sc., 
Botanical Laboratory , University of Liverpool. 
[With Text-Figs. 19—25.] 
OLLIQU AY A ODORIFERA is a component of the bushy 
sclerophyllous woodland which occurs on the slopes of the 
Chilian Andes at heights between one and two thousand feet. An 
account of the anatomy of the tribe Hippomaneae of the Euphor- 
biacese is given in an inaugural dissertation for the degree of doctor, 
accepted by the Faculty of Philosophy in Munich in 1897. The 
author, Herbert, describes the leaf and axis of C. odorifera, but his 
descriptions were, as he says, based upon the examination of 
herbarium material to which he had access. The leaf is large and 
ovate with a serrate margin, having at the apex of each tooth a 
prominent, black, almost spherical “gland.” These glands, which 
form the most characteristic feature of the leaf, are neither 
mentioned by Herbert in his dissertation, nor by Solereder in his 
Systematic Anatomy of the Dicotyledons (English Edition, 1908). 
It should be mentioned, however, that the latter author relies to a 
considerable extent on the former for information as to the anatomy 
of the Hippomaneae. 
While investigating the structure of these glands the writer 
made an examination of the other portions of the leaf of C. odorifera, 
and as his conclusions did not agree in all respects with those of 
Herbert it seemed advisable to publish them. 
The material investigated was taken from a plant, raised in a 
greenhouse by Mr. G. F. Scott-Elliott from seed collected by that 
gentleman in Chili. A large well-grown twig which was preserved 
in spirit was used. This twig contained several large full-grown 
leaves together with numerous smaller ones. Microtome sections 
of most of these leaves were cut and the following description is 
based on them. 
Epidermis. —There is very little difference between the cells of 
the upper epidermis and those of the lower in shape, size or thick¬ 
ness of cuticle. The former, are, however, not uniform, being 
larger near the base of the leaf and also near the middle. Over the 
median nerve they lose their typical structure and become collen- 
chymatous. These collenchymatous epidermal cells overlie a bundle 
