144 
M. Benson. 
Sphenophyllales. The evidence in favour of this “ reduction 
hypothesis ” is still very inadequate, although Miss Sykes’s work 
on the “ Sporangium-bearing Organs of Lycopodium 1 ” has shown 
the wide range of form and structure that the “ sporange” exhibits. 
For the sake of comparison with her results it may be interesting 
to refer to two Palaeozoic types of Lycopodinean “ sporange ” which 
unfortunately still await description. I append diagrammatic figures. 
Fig. 25. Diagrams of halves of tangential sections of a micro- and 
megasporophyll respectively, x 30. Royal Holloway College Collection, C.N., 
266 - 4 and 192n. Both specimens are referred to as Lepidostrobus Mazocarpon. 
Fig. 25 is from two specimens met with in the Burntisland 
rock, a and a 1 represent halves of tangential sections of a micro- 
and a megasporophyll of what I assume to be a species of 
Lepidostrobus. They are associated with the branching stems of 
LepidodeudronPettycurense, which has recently been shortly described 
by Mr. Kidston 2 as showing a solid protostele. Owing to the resem¬ 
blance in some respects which these sporophylls show to Mazocarpon 
1 will refer to them here as Lepidostrobus Mazocarpon. They are 
interesting not only because of the large amount of sterile tissue, 
but because of its character. A wide layer abutting on the sporo- 
genous regions consists of elongated tapetal cells. The divergence 
of these cells leads to a split which may be seen in the median plane 
of the megasporophyll. This is somewhat suggestive of a multiple 
origin of the sporogenous region. 
1 Sykes, New Phyt., 1908, Vol. VII., p. 41. 
2 Kidston, Proc. R. S, Edin., Vol. XXVII. 
