2 I 2 
W. Stiles 
on either side it is produced into a membranous lamina, to the 
lower side of which are attached the two microsporangia, one on 
each side of the sporophyll axis. The cone is surrounded at the 
base by an involucre-like ring of acute scales, each containing a 
vascular bundle and resin-canal similar to those of the leaf. 
The axis of the cone contains a ring of small collateral 
bundles, the most internal elements being frequently separated 
from the rest of the xylem. The presence of these few scattered 
tracheids points to the occurrence of centripetal xylem. As usual 
there is a resin-canal outside the phloem of each bundle. 
The bundle of the sporophyll is given off from one of the axial 
bundles by division of the latter in a tangential plane. It is very 
small and consists largely of phloem, only a single tracheid or two, 
or three tracheids at the most, being present in the pedicel. The 
bundle is accompanied by a resin-canal on the under or phloem 
side. The wall of the sporangium is one cell wide in the mature 
cone except on the outside of the latter, and the vertical cell walls 
are rather thick (Fig. 29). The wall is strikingly similar to that of 
Fig. 29. Vertical section through a male sporophyll, not median, x 80. d, 
point where the section cuts the line of dehiscence of the sporangium ; 
s, large cells with deeply-staining contents between the outer wall and 
the tapetum. 
Araucaria Rulei, figured by Seward and Ford 1 , while the large 
secretory cells of the latter between the wall and tapetum, strongly 
suggest the large cells with deeply staining contents occupying 
a similar position in Saxegothrea (Fig. 22s). Dehiscence of the 
sporangium takes place transversely by means of a slit on the 
lower surface; its position is cut across transversely in Fig. 29 at 
the point d. On the splitting of the wall the part on either side of 
the slit rolls back so that at first sight the sporophyll appears to 
end in a large reflexed tip as described by Lindley 2 . This writer’s 
1 Seward, A. C. and Ford, Miss, S. O. (27), p. 356. Fig. 22 h. 
2 Lindley, J. (18). 
