2 14 
W. Stiles. 
It is noteworthy that the projecting apex of the nucellus is more 
prominent than in Araucaria; its great width and uneven surface 
are striking features in which it resembles the stigma of an 
Angiosperm. 1 
The material examined by me had reached a stage in which a 
cavity, possibly the embryo-sac, had appeared in the middle of the 
nucellus (Fig. 31c). The cells in its neighbourhood are grouped 
more or less concentrically round it, while in the micropyle they are 
Fig. 31. Longitudinal median section through the ovule, o, outer integument; 
i, inner integument; n, nucellus; st., nucellus extruded; c, central 
cavity. 
somewhat lengthened longitudinally and often shew remarkable 
thickening bands on their walls; the cells of the stigma-like 
expansion are not so regularly arranged. It is hoped to continue 
the study of the megasporangium in the future when more material 
may be available. 
After fertilisation the scale swells up and becomes fleshy, and, 
.by the fusion of the scales of the cone, a globular fleshy mass is 
produced, containing the shiny pale brown seeds. 
1 M. Tison (l.c.) finds that the projecting apex is covered by a 
sticky secretion and presumably acts as a stigma ; and also 
that it continues to grow in size for about two months after 
pollination. 
