W. Stiles. 
2 18 
the scale, while in these species of Araucaria the first bifurcation 
takes place in the cortex of the cone axis ; and, secondly, the bundles 
of the upper row do not bend round to the sporangium in A. Cookei 
and A. Rulei as they do in Saxegothaea. The vascular system in A. 
Bidwillii and A. imbricata is developed less like that in Saxegothaea, 
but in the former Worsdell 1 observed strands of phloem-like tissue 
entering the chalazal end of the ovule. The method of branching of 
the bundles in Agathis 2 is different, in that the single bundle divides 
into a row of about twelve, and then the median one of these gives 
rise to the upper row. Transfusion tissue is prominent in both Saxe¬ 
gothaea and the Araucarieae. It is of importance to note that in 
both cases also a portion of the nucellar tissue projects through the 
micropyle. On the other hand, Saxegothaea differs from the 
Araucarieae in having the ovule inverted 3 ; it further differs from 
Araucarieae in not having the sporangium immersed in the tissues 
of the scale, and in the absence of a ligule, in which respects it 
agrees with Agathis. 
Saxegothaea differs from all other Podocarpeae except Micro- 
cachrys tetragona in having its female sporophylls arranged in a cone. 
The arrangement of the vascular bundles in the scale of Micro- 
cachrys 4 is not unlike that of Saxegothaea, but more closely resembles 
Araucaria Bidwillii. The mode of attachment of the ovule in 
Microcachrys is also not unlike that of Saxegothaea. Transfusion 
tissue is also present in the female sporophyll of Microcachrys, but 
is wholly lateral in position. 
The presence of only one resin-canal in the female scale of 
Saxegothaea is interesting, as in all other parts of the plant a single 
canal accompanies each primary vascular bundle on the phloem 
side. If the scale of Saxegothaea were a double leaf-structure it 
would not be unreasonable to expect to find two resin-canals, one 
on the phloem side of each bundle. The state of things found 
apparently indicates that the scale of Saxegothaea is what it at first 
sight appears to be: a simple sporophyll bearing on its upper 
surface a single megasporangium. 
1 Worsdell, W. C. (31), p. 532. 
3 Seward, A. C. and Ford, Miss S. O. (27), p. 360. 
3 M. Tison finds that the ovule of Saxegothaea is at first perpen¬ 
dicular to the scale, but my material was not young enough 
to shew this. 
3 Worsdell, W. C. (31), p. 537. 
