James Small. 
258 
With the same touch the anther tube is stimulated to bend towards 
the touch, and this irritability remains during anthesis. The style 
is also irritable, and with the first touch after exsertion moves as in 
Arctotis, but further touches have little or no effect for at least half 
an hour unless the same side of the style is touched, when a slight 
movement in the same direction as the first ensues. 
The parents of Gazania splendens X are not known, but are 
said to be two of the three species— G.ringens, Gaertn., G.pavonia, 
R.Br. and G. uniflora, Sims. 
It is noteworthy that all these three genera belong to the 
Arctotideae, while three of the genera ( Mimulus , Torenia and 
Glossostigma) in the other families belong to the Gratioleae of the 
Scrophulariaceae, and that these other families themselves all belong 
to Sections D and E of the Tubiflorae (Engler). The phenomenon 
seems to be restricted to a few narrow circles of affinities, and is 
not wide spread as in the case of irritability of the stamens. 
C. Irritability of the Stamens. 
The basis of the present section is the following list, Table VIII, 
(pp. 260-266) of the species on the irritability or non-irritability of 
which observations have been made. The tribes are arranged accord¬ 
ing to their origin as shown by the structure of the pollen-presentation 
mechanism, and the sub-tribes are those of Bentham revised as 
before. The recognised affinities of the genera are indicated roughly 
by the numbers, which are those of the Genera Plantarum, except 
in cases where Bentham sunk a genus which is now upheld, e.g., 
Leptosyne, when a letter is prefixed to the Genera Plantarum 
number of the containing genus. 
Previous authors have not as a rule recorded the exact type 
of irritability, and those species not observed in the present 
investigation are indicated thus :— 
(Cov) — irritability of the stamens recorded by Covolo; 
(K) = „ 
(Kb) = 
(Kn) = 
(H) = „ 
(M) = „ 
(L) = ,, 
(U) = „ 
(J) = „ 
(OJ)— absence of irritability recorded by Juel 
Koelreuter; 
Kabsch; 
Knuth; 
Haberlandt; 
Meehan; 
Linsbauer; 
Unger; 
J uel; 
