Origin and Development of the Compositee. 259 
The majority of the species have been examined by the writer 
and the types of irritability are distinguished thus:— 
Types of irritability : 
A : pollen presented on touching; no lateral movement of the 
anther tube, 
B : pollen presented on touching; lateral movement indefinite 
in direction, 
C : pollen presented on touching; lateral movement towards 
the touch, 
C': pollen presented on touching; lateral movement towards 
the centre of the capitulum, 
C e : pollen presented on touching with more or less explosive 
rapidity, 
O : no irritability observed when touched. 
The degree of irritability is noted as slight when it requires 
careful observation, and various other degrees are noted. Various 
precautions are necessary for the successful observation of the 
phenomenon, for it was not without reason that Camerarius 
remarked of the observation of the movement that “ Verum 
enimvero patientia.” The general method used in the present 
investigation was to examine the flowers on a hot, sunny day 
just after anthesis had commenced, and before an insect had 
visited the floret, by touching the filaments or anther tube gently 
with the point of a needle, and by observing the effect with the 
aid of a watchmaker’s eyeglass. The latter was found very con¬ 
venient as it left both hands free for manipulation. For the details 
of the precautions readers are referred to the previous paper (49). 
The list includes 360 species and varieties, 253 or 70% of 
which show irritability ; all the tribes in the family are represented 
and 45 of the 64 subtribes, so that the list is sufficiently repre¬ 
sentative to form the basis of some phylogenetic suggestions. 
Notes on Table VIII. 
1. In all these cases (Tussilagininae) the critical stage, with 
the anther tube open and the style not completely exserted, is 
very short. 
2. In Doronicum Pardalianches the style has frequently been 
observed to be bent by pressure against the top of the anther tube 
3. Cineraria Moorei is a garden hybrid between C. Heritieri 
and a form of C. ententes. \ 
4. Cineraria stellata is a florists’ variety obtained from 
Messrs. Dobbie and Sons. 
