Lepidocarpon. 31 
functional macrospore and provided with an integument which 
grows up from the body of the sporophyll. This integument, 
together with the sporophyll, completely envelopes the sporangium. 
A number of cones have been examined by Dr. Scott, viz.—(1) 
Younger stages of macrosporangiate strobili having the radially 
elongated sporangia seated on the sporophylls precisely as in 
Lepidostrobus . The only peculiarity of this stage, and it is an 
important one, resides in the fact that only a single functional 
macrospore is present in each sporangium, accompanied by the 
other three members of the tetrad in an abortive condition. The 
organisation of the cone is wholly that of a Lepidostrobus , for which 
it may readily be mistaken. (2) An older cone, differing from the 
foregoing in the presence of integumented in addition to naked 
sporangia. The integument grows up from the flanks of the 
sporophyll, meeting above the sporangium. The line of closure of 
the integument is a long crevice running the whole length of the 
upper surface of the sporangium. This slit has been compared to 
a micropyle. A point of interest and perhaps of significance is the 
existence of numerous non-integumented sporangia, having the 
appearance of maturity in their tissues. Dr. Scott regards them as 
arrested stages rather than merely young ones, though they may 
be taken as representing in essentials the young condition hardened 
and matured, and thus better fitted for preservation than specimens 
in the normal course of development. The meaning of this frequent 
arrest is not easy to understand, but it seems futile to speculate on 
the matter till more developmental data are available. (3) a 
microsporangiate cone from the same block as one of the younger 
macrosporangiate strobili, and associated with detached integu- 
mented and non-integumented sporangia. This cone, whose 
reference to the same species seems more than probable, has a 
peculiar interest in that its sporangia are integumented in the 
same way as are the macrosporangia, though the enclosure is not 
so complete. The homology of the integuments in the two cases 
seems evident. 
In addition to these cones there have been examined a large 
number of detached sporophylls with integumented and non- 
integumented macrosporangia. In a few instances among the 
former (the “ seeds ”) the macrospore contains a prothallium. The 
* 
prothallium shows peculiarities suggestive of archegonia and 
possibly of an embryo, but the data are not sufficiently precise for 
a certain determination. 
