Edith R. Saunders. 
58 
variations in amount of light and moisture are probably to be 
reckoned. Whether other factors are also concerned, and how far 
they may be opposed in their action, further experiment must 
decide. These conclusions are in harmony with the view held by 
Peyritsch 1 and others that the condition of peloria is also to some 
extent affected, if in some cases not actually induced by the 
stimulus of changes in the environment. So far as I am aware, 
however, no critical experiments on this point have as yet been 
carried out. 2 
It is further evident that the Jieptandra form is related to the 
type as recessive to dominant. Plants derived from the original 
Jieptandra parent but resembling the type—hence evidently cross¬ 
breds—invariably gave a mixed offspring, the numbers approximating 
to the ratio 3 Type : 1 Heptandra, or 1 Type : 1 Heptandra, according 
as they were fertilised with their own or sister pollen, or were 
crossed back with the heptandra form. Details of the breedings are 
as follows:— 
(1) . Nine of the putative crossbreds derived from the original 
Jieptandra plant were self-fertilised, and one of these individuals 
was also intercrossed with a sister plant. In all ten families the 
offspring were mixed; the total numbers obtained were 183 Type 
(including one plant showing slight abnormality in the first four 
flowers) and 49 Jieptandra (including ten plants with some normal 
or nearly normal flowers), where, on the basis of a 3: 1 ratio, we 
should expect 174 Type and 58 Jieptandra. 
(2) . Three of the crossbreds, of which two were among those 
self-fertilised in the preceding experiment, were crossed back with 
various Jieptandra individuals. The four resulting families were all 
mixed, the totals obtained being 69 type (including seven individuals 
with a few abnormal flowers) and 59 Jieptandra (including six 
individuals with a few normal flowers) where we should have 
expected equality. 
1 Untersuch. iiber die Aetiologie pelorischer Bliithenbildungen. 
Denkschr. d. k. Akad. Wien. Bd. XXXVIII. 
s Gallardo’s observation on the higher proportion of peloric 
individuals occurring among a batch of foxgloves grown 
under favourable conditions as regards space and illumination, 
as compared with another batch of similar origin, raised 
simultaneously, but less exposed to the sun and more crowded 
together, cannot be used as evidence in this connection, since 
the peloric parents were not apparently protected from 
crossing. Had intercrossing with type plants been prevented, 
all the plants in both batches would presumably have been 
peloric since peloria in the foxglove comes true from seed. 
(See Gallardo. Notes morphologiques et statisques sur 
quelques Anomalies hereditaires de la Digitale. Rev. gen. de 
Botanique XIII., 1901, p. 163). 
