Inheritance of Mutation in the Common Foxglove. 61 
been used as a parent in the case of the other four families, but 
instead of being self-fertilised they were each crossed back with two 
different heptandra individuals. Each again gave a mixture of hairy 
and smooth, but the proportion of smooth was now greatly 
diminished, as shown below. 
Comparison of the six mixed families derived from the two Fj 
crossbreds (plants A and B) showed the following results:— 
Hairy. 
Smooth. 
Plant A X self gave 
6 
23 
Plant B x self gave 
5 
13 
Totals 
11 
36 
Hairy. 
Smooth. 
Plant A x heptandra plant 1 gave 
29 
16 
X heptandra plant 2 gave 
11 
12 
Plant B x heptandra plant 3 gave 
13 
9 
X heptandra plant 4 gave 
17 
7 
Totals 
70 
44 
It is somewhat surprising, in view of the relation which 
experiments in other cases have shown commonly exists between 
the hairy and the smooth condition that the two sporting individuals 
in this case should have produced an excess of smooth plants on 
self-fertilisation. For an explanation of this result we must await 
the evidence from later generations. 
The observations made in regard to the inheritance of flower 
colour are in entire accord, so far as they go, with the facts already 
published last year by Keeble, Pellew and Jones, 1 viz: 
(1) . That in all individuals whether white or coloured the 
lower lip is spotted, the spots being of various sizes. 
It was noticed that in some families the spot area was occupied 
by large irregular blotches of colour, the appearance suggesting 
that a number of spots of ordinary size had become confluent. In 
one family this blotching was observed in ten out of the twenty- 
eight individuals recorded ; in the remaining two families in whicq 
this peculiarity was seen the proportion was very much fewer. It 
is hoped to obtain evidence in regard to the inheritance of this 
character in the next generation. 
(2) . That among white flowered plants there are two kinds of 
individuals:— 
(a) White-flowered plants with yellowish-green spots. 
(b) White-flowered plants with red spots. 
1 The Inheritance of peloria and flower colour in Foxgloves 
[Digitalis purpurea). New Phytologist, Vol. IX., 1910. 
