Methods of Reproduction in the Saprolegniacecz. 177 
older hyphae thus become “ torulated ” the whole mycelium has the 
appearance of being punctated. The swellings (or gemmae) are 
either club-shaped, pear-shaped, cylindrical, or sometimes irregularly 
shaped, and of very different size. On germination the gemmae give 
rise to secondary gemmae, secondary sporocysts or to oocysts. The 
sporocysts are of two kinds. The first kind resemble those of other 
species of Saprolegnia, as already described. The second kind 
occur later and are formed in a row one behind the other, or 
variously arranged according to the arrangement of the “ swellings 
(chain gemmae) from which they are formed. 
The terminal sporocyst of such a row discharges by a terminal 
process, the other by a lateral opening (figs. 11 and 14, PI. I., 
Vol. IX). 
Oocysts are formed from the swellings on the hyphae and in 
the older filaments often differ very much in shape and size, some¬ 
times in a row one behind the other and, more rarely, completely 
isolated. The oocysts remain firmly attached to the mycelial 
filament until the oospores are mature; they have a smooth, 
coloured, clearly defined wall, not punctated, and contain many 
oospores. 
Antherocyst usually absent, but when present arises either 
from the end of an adjacent branch of the same hypha (androgynous), 
or from a separate branch of the mycelium (diklinous). 
Oospores, many in each oocyst, spherical or sometimes unevenly 
polygonal, smooth membrane, 14— 22g in diameter. Resting period, 
ten days ; on germination give rise to a mycelium directly or to a 
sporocyst. Gemmae of different shapes formed from the swellings 
on the “torulose” hyphae ; on germination they give rise to a germ- 
tube or a sporocyst. 
The foregoing account agrees in all essential features with 
the present species, the only point of difference being the duration 
of the resting period of the oospores. In the case of the present 
species only one case of germination has been observed (p. 172, 
fig. 16). The numerous oospores produced in the cultures showed 
no tendency to germinate even at the end of a month. 
With the exception of this slight difference, the species agree so 
very closely that there seems to be no doubt they are identical. 
Conclusion. 
The present species, did not show the great variety of 
methods of asexual reproduction shown in the last species, 
