r 18 On the Megaspore of Lepidostrobus foliaceus. 
the microsporangiate cone of this plant in order to understand the 
details of its structure. The original slide is in the Natural History 
Museum, C.N. 1614. This is the type specimen on which Mr. 
Fig. 25. Photograph of 5 (Plate VIII.) Sporangia containing megaspores, 
one with the characteristic bract of Lepidostrobus foliaceus. 
7ii., megaspore. 7iig., megasporangium, b., bract. 
Maslen founded his species Lepidostrobus foliaceus. The section 
is a fairly radial one through the microsporangiate part of the cone. 
It is not quite median but it clearly shows the elongated form of the 
sporangia seated on the sporophylls with their leaf-like laminae 
from which the specific name was taken. 
We have been trying for the last year to sort out the different 
megaspores of Lepidostrobus, so I looked through our camera lucida 
drawings of megaspores to match those found in the sporangia. 
The result was, that the only ones at all corresponding in size 
and texture were my other new megaspores Triletes diabolicus. I 
searched carefully to see if 1 could find any sign of the curious 
appendage, but without success, till when 1778 was reached I had 
the good fortune to find a sporangium containing a typical Triletes 
diabolicus. 
Since then 1 have been through every slide of Lepidostrobus 
foliaceus and have found that 25 out of 36 show the Triletes. 1 now 
thought if the type cone of Williamson were examined and showed 
Triletes diabolicus in the cone the evidence would be conclusive. 
The cone, however, so far as shown, is purely microsporangiate, 
but the section shows a very fine specimen of Triletes diabolicus 
completely enclosed in a sporangium, identical in appearance with 
those containing the microspores, so that 1 think there is little doubt 
that the two new megaspores are really one and the same, that the 
Williamson cone must in future be considered as a heterosporous not 
a homosporous one and that Triletes diabolicus must disappear and 
become the megaspore of Lepidostrobus foliaceus , 
