Recent Advances in the Study of Heredity. 
33 3 
coupling occurs in such a way that the gross, character is associated 
with maleness, and the lacticolor character with femaleness. 
3. In the 3 s which are colour heterozygotes no such coupling 
occurs. 
4. There is a preferential mating amongst the gametes, such 
that a union can only take place between 3 - and ? -bearing 
gametes. 
5. Dominance is determined by the female parent. 
Bateson has put forward a much simpler explanation of these 
phenomena, which, like his theory in explanation of Correns’ results 
with Bryonia , starts from the assumption that femaleness is 
dominant. His theory is as follows :— 
1. The female is heterozygous in regard to sex, femaleness 
being dominant; the male is a homozygous recessive. 
2. In the gonads of the hybrid in which the two dominant 
factors femaleness and the gross, character co-exist there is spurious 
allelomorphism between these two dominant factors, that is to say 
each gamete may bear one of these factors, but not both. The 
following analyses of the matings shows that the actual results follow 
from this theory:— 
1. lact. ? X gross. $ 
RR ? 3 I DD 3 3 
F 1 gross. 3 
DR 3 3 
2 . 
X 
2 gross. 3 
DR 3 3 
DD $ 3 
lact. ? 
RR? 3 
1 gross. ? 
DR ? 3 
X F, gross. 3 
DR 3 3 
1 gross. 3 
DR 3 3 
1 gross. ? 
DR ? $ 
Fj gross. ? 
X 
1 lact. 3 
RR 3 3 
lact. 3 
gross. ? 
DR ? 3 
composition 
gametes 
DR ? 3 
D 3 
R? 
\ 
DR 3 3 
D 3 
Ref 
1 
gross. 3 
DR 3 $ 
't Q 
RR ? 3 s 
1 lact. ? 
RR ? 3 
lact. ? 
RR ? 3 
The following Tables exhibit the above mating in a form which 
I find is more easily intelligible ;— 
