Origin of QL nothera Lamarckiana De Vries. 237 
What we desire is further information on the composition of 
the cultures of Carter and Company through the discovery of other 
herbarium material of about the same date (1862) as that of the 
sheet in the Gray Herbarium. It will be strange if Dr. Gray proves 
to the only botanist who preserved specimens of the “ Lamarckiana ” 
placed on the market by Carter and Company. There should be 
made a persistent search through herbaria to bring forward any 
specimen that may throw light upon the problem. This is a matter 
in which it would seem that the British botanists can render a 
great service. 
Another feature of the problem concerns the development of 
some remarkable CEnothera floras in parts of England composed in 
greater part of 0. Lamarckiana or variants from this type. 
It is surprising how common have become Lamarckiana -like 
forms in England. During the past three years several English 
botanists have kindly replied to my request for seed of broad-leaved 
forms of CEnothera biennis with green stems bearing red papillae at 
the base of long hairs. From the seed sent to me I have grown 
eleven different cultures in the hope of finding a type of biennis 
which in the past I have greatly desired as a parent for a cross 
with O. graudi/iora. All of these cultures have proved to be forms 
essentially Lamarckiana as to habit, foliage and stem coloration, 
but with smaller flowers than is usual for De Vries’s plant. They 
correspond very closely and some ol them are indistinguishable 
from the small-flowered races of Lamarckiana which I have 
differentiated from material of De Vries.' They were not at all 
the forms of biennis that I hoped to obtain, and could not be used 
in my experimental work. 
The type of biennis that I wish should agree closely in 
morphology with the biennis of the sand dunes of Holland, but it 
should have the stem coloration characteristic of Lamarckiana, i.e., 
the green portions of the stems should be punctate with red papillae 
at the base of the long hairs. The Dutch biennis has as far as we 
know a clear green stem above, but it would not be surprising if a 
form should be found with the stem coloration of Lamarckiana, 
since races of American biennis occur differing only in the presence 
or absence of red coloration in the stem papillae. Seeds of a type 
agreeing with the Dutch biennis have been sent to me from the 
botanical garden of Cambridge University and presumably the plant 
See Davis, American Naturalist, vol. 46, p. 3S3 1912. 
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