Origin of CEnothera Lamarckiana De Vries. 241 
Thus Don 1 described in 1832 “(E. biennis” from “sand banks on 
the coast of Lancashire ” and noted that it “ is common in gardens 
and often escapes from thence into rich waste ground,” but the 
statements “ flowers large ” and “ delicately fragrant ” are relative 
matters and we cannot be certain from this or other parts of the 
account whether the plants were nearest to Lamarckiana De Vries 
or to forms of biennis such as that from Holland (the biennis of 
Linnaeus). Lindley 2 in 1833 gave an account of 11 CEnothera 
biennis var. graiulijlora ” and stated it to be “ not uncommon in 
gardens.” The figure of this plant shows petals drawn about 
3.5-4 cm. long, as large as those of Lamarckiana De Vries, but a 
basal or perhaps a rosette leaf is too narrow for Lamarckiana, nor 
will it do for the Dutch biennis or for grandiflora. The bracts are 
figured broad at the base and sessile as in both Lamarckiana and 
biennis, but the inflorescence with long internodes is not as in those 
species but is more like that of grandiflora ; the bracts, however, 
are not those of grandiflora. These contradictions are most puzzling, 
but may there not be specimens preserved at this period which will 
clear the obscurity. 
I am aware that the CEnothera floras in other parts of England 
offer opportunities for investigation that should not be overlooked 
in the study of the problems presented in this paper. I have, 
however, confined my suggestions to the remarkable assemblage on 
the sand hills of Lancashire for the reason that this region seems 
to be best known and to have been under observation for the 
longest period. 
] Don, George. A General System of Gardening and Botany, vol. 2, 
p. 685, 1832. 
1 Lindley, John. CEnothera biennis ; var. grandiflora. Edwards’s Botanical 
Register, vol. 6, p. 1604, 1833. 
University of Pennsylvania. 
March, 1913. 
