The Classification of Conifers. 255 
nuclei, rosette, suspensor and embryo tiers); male and female 
nuclei very unequal in size at the time of fertilization. - Pinacece. 
III. i. Archegonia as in family; male gametopliyte with two 
prothallial cells, not persistent; male cells nearly equal; pro-embryo 
forming eight free nuclei at base of archegonium, before wall- 
formation ; four tiers of (a) four free nuclei, (b) four rosette cells, ( c ) 
four suspensor cells, and (d) four embryo-cells Pinacece: Abietoidece. 
III. ii. Archegonia as in family ; male gametopliyte with no 
prothallial cells; two nearly equal male nuclei lying free in the tube 
cytoplasm ; pro-embryo forming more than eight free nuclei at base 
of archegonium; four tiers formed as in III. i. but terminal tier 
dividing to form a group of about sixteen cells before elongation of 
the suspensors. .... Pinacece: Sciadopitoidece. 
IV. Archegonia apical or lateral, more or less united in one 
or more groups or complexes, with a common jacket-layer (where 
present); male gametopliyte without prothallial cells; male cells 
exactly equal and clearly defined, at first hemispherical, and both 
functional as a rule; pro-embryo with not more than eight free 
nuclei before wall-formation, otherwise differing widely in the three 
sub-families; male and female nuclei not differing widely in size at 
the time of fertilization. ..... Cupressacece, 
IV. i. Archegonia in a single apical complex, surrounded by 
a common jacket-layer (in one genus some vegetative cells are 
included in the centre of the complex) ; male gametopliyte as in 
family ; pro-embryo forming eight free nuclei before wall-formation, 
occupying only a small part of the archegonium at its base; mature 
pro-embryo usually of three tiers or regions, less regular in arrange¬ 
ment than in III, commonly with one of the cells of the embryo 
region in advance of the others ; male nucleus distinctly smaller than 
female at the time of fertilization. - Cupressacece : Cnpressoidece. 
IV. ii. Archegonia in one or (usually) more lateral groups, 
each with traces of a common jacket-layer; apex of prothallus 
pointed; male gametopliyte as in family; pro-embryo forming four 
free nuclei before wall-formation, filling the archegonium, and 
dividing up into several groups of two or three cells, of which one 
becomes the suspensor and one the embryo initial; male and 
female nuclei equal (or almost equal) in size at the time of 
fertilization. ..... Cupressacece: Callitroidece. 
IV. iii. Archegonia in several lateral groups; apex of pro¬ 
thallus pointed; male gametopliyte as in family; pro-embryo 
becoming walled at first division ; mature pro-embryo filling the 
archegonium, and consisting of about five cells, of which the 
