AMBLYPODA. 
181 
hoofed Mammals, and maintains itself with great persistency in both of 
them. It is essentially a primitive character, the alternating position being 
usual in the cold-blooded Vertehrata, and is the persistence of the oblique 
relation of the original divergent branching rays, to which digits have been 
traced. In the Proboscidia and Hymcoidea, the elements of the two rows 
assume an opposite and longitudinal relation. The structure of the fore 
foot in the Amblypoda appears to be about equally related to that of the 
Proboscidia^ the Perissodactyla, and the Artiodactyla. 
In the cubito-carpal articulation, the resemblance is again to the 
Proboscidia in the relatively large proportion of it belonging to the ulna, 
and the consequent lateral position of the lajtter bone. In this respect, it 
differs much more from the other two living orders of hoofed Mammals, 
although here again the lower forms of both resemble the Amblypoda more 
than do the higher forms. As is well known, both of the hoofed orders 
display a constantly diminishing extent of the ulno-carpal articulation, and 
increase of the radio-carpal, until, in the Horse and Ox, the idna becomes 
a mere splint attached to the radius. 
The relationships indicated by the brain are to the lissence})halous 
orders Chiroptera, Insectivora, and Edentata. As an ungulate order, the 
Amblypoda are distinguished from the first two, were other characters 
wanting. We may here notice, however, some curious resemblances 
between the forms of the teeth and lower jaw of Coryphodon and some 
Insectivora, and the still more curious resemblance betAveen the tibio-tarsal 
articulation in the order and that of the cotemporary Creodont allies of 
the Insectivora. Comparison with the ungulate forms of Edentata only is 
necessary, and from these the enamel sheathing of the teeth separates the 
Amblypoda at once. The small size of the brain doubtless relates these 
animals to the other Eocene Ungulata, described by Lartet, still more 
nearly than to existing Lissencepliala. In the small size and smoothness of 
the hemispheres, and relatively large development of the optic and olfactory 
lobes, the brain of the Amblypoda more nearly resembles that of the 
Creodonta than that of any division of recent animals. The resemblance 
between the brains of Amblypoda and those of the Carnivorous Oxycena 
and Arctocyon {fide Gervais), is so great as to testify to a similar degree of 
