86 
DR. C. E. BEEVOR AND MR. V. HORSLEY ON THE EXCITABLE 
of fibres in the capsules, still less differentiation would be found. This is obviously 
shown in Table V., but on inspection, the representation of even flexion of the small 
toes is consistently (every group) in front of that of extension. To a minor degree 
and witlr less consistency (4 of 6 groups) the same holds for the hallux. 
Relative Amount or Degree of Representation of Extension compared to Flexion. 
This subject may not be without interest to those who have, from excitation 
experiments, attempted to gauge the relative amount of functional activity in the 
cortex. 
The facts bearing upon this point obtained from the records of hinge joint move¬ 
ments are so clear and unmistakable that they are of value. 
The following are the proportional rates of frequency as observed in the capsule. 
Flexion. 
Extension. 
Remarks. 
Elbow .... 
Fingers 
.38 
38 
IG 
17 
I Joints whose usual movement is powerful flexion 
Thumb . . . 
22 
9 
Joint the flexion of w'hich is usually carefully regulated ' 
by extension 
Small toes 
1.5 
26 
Joints very frequently spread in extension for standing 
Hallux . . . 
16 
17 
Joint used as a thumb as well as for support 1 
The bearing of these facts obviously requires no further comment, and the more so 
as they are but confirmatory of the results we obtained in our analysis of the cortical 
representation of these parts. 
Conclusions. 
As the foregoing paper is but a collection of detailed facts we can only offer as 
conclusions a few bare statements. 
1. The fibres of the excitable portion of the internal capsule are arranged from 
before back, in the same order as the foci of representation in the excitable part of 
the cortex, taken along lines drawn at right angles to the direction of the fissure of 
Kolando. 
2. This applies to the great divisions of the body, i.e., head, upper limb, &c. 
3. The arrangement of the fibres for the different segments of each great division is 
likewise that which prevails in the cortex. 
4. Similarly the character of each movement is represented in the capsule as in the 
cortex. 
