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III. Mathematical Contributions to the Theory of Evolution. —VIII. On the Inheri¬ 
tance of Charade,not capable of Exact Quantitative Measurement .—Part I. 
Introductory. Part II. On the Inheritance of Coat-colour in Horses. Part III. 
On the Inheritance of Eye-colour in Man. 
By Karl Pearson, F.R.S., with the assistance of Alice Lee, D.Sc., University 
College , london. 
Received August 5,—Read November 16, 1899 ; withdrawn, rewritten, and again l’eceived 
Maixh 5, 1900. 
Contents. 
Part I.— Introductory. 
Page 
§ 1. General Nature of the Problem and Assumptions upon which it can be solved. 80 
§ 2. Determination of the Mean Value of the Characters and the Ratio of the Variability of 
Correlated Characters. 81 
§ 3. Determination of the Probable Errors and Error Correlations of ail the quantities involved . 82 
§ 4. On the Construction of Normal Scales for Characters not capable a priori of exact 
quantitative Measurement. 87 
§ 5. On Blended and Exclusive Inheritances. 88 
Part II .—On Coat-colour Inheritance in the Thoroughbred Horse. 
§ 6. On the Extraction and Reduction of the Data. 92 
§ 7. On the Mean Coat-colour of Thoroughbred Horses. 93 
§ 8. On the Relative Variability of Sex and Generation. 94 
§ 9. On the Inheritance of Colour in Thoroughbred Horses. 98 
(«) Direct Line, First Degree. ( b ) Direct Line, Second Degree. (c) Collateral 
Inheritance. General Conclusions. 
Part III .—On Eye-colour Inheritance in Man. 
§10. On the Extraction and Reduction of the Data. 102 
§ 11. On the Mean Colour, having regard to Sex and Generation. 104 
§ 12. On the Relative Variability of Sex and Generation .. . 109 
§13. On the Inheritance of Eye-colour in Man. .. 113 
(ft) Assortative Mating. (b) Collateral Heredity, First Degree. (c) Collateral 
Heredity, Second Degree, (cl) Direct Heredity, First Degree, (e) Direct 
Heredity, Second Degree. (/) On Exclusive Inheritance. 
§14. General Conclusions.. 119 
Appendix I. Tables of. Coat-colour in Horses. 122 
II. Tables of Eye-colour in Man. 138 
Note I. Inheritance of Temper and Artistic Instinct. 147 
Note II. On the Correlation of Fertility and Eye-colour.148 
YOL. CXCV.—A 264. 
29.10.1900 
