November, 1999 
SCAMIT Newsletter 
Vol. 18, No. 7 
at Eniwetok Atoll of Marshall Islands, in 1971 
at New Caledonia, and in 1975 during the 
Alpha Helix East Asia Bioluminescence 
Expedition to Moluccas. 
In the 50’s Arthur completed his studies of 
copepods collected during his sabbatical leave 
to West Africa. During the 60’s he published 
mostly on the copepods collected from West 
Indies and Nosy Be. In the 70’s there was a 
gradual shift in his studies from the Caribbean 
Sea and western Indian Ocean to the copepods 
living in association with various invertebrates 
occurring in Eniwetok Island, Mollucas, and 
New Caledonia. Arthur began in the 80’s to 
publish his works on the copepods collected in 
the water around deep-sea hydrothermal vents, 
which had been discovered in the late 70’s, as 
well as the copepods associated with vent 
animals. 
Arthur’s greatest contribution to copepodology 
is his discovery and description of many 
symbiotic copepods which occur in association 
with a diversity of marine animals, ranging 
from primitive sponges to more specialized 
marine mammals. In his half century (1941- 
1999) of work, he published more than 250 
papers and monographs on the symbiotic 
copepods, described no less than 650 new 
species and created more than 140 new genera 
and 16 new families. An exact number of 
species and genera of copepods new to science 
cannot be determined at this time because there 
are manuscripts by Arthur either in press or 
submitted for publication. In the history of 
copepodology, no copepod taxonomist has 
been as productive. 
More than a wonderfully effective taxonomist, 
Arthur was a distinguished teacher as well as 
an excellent editor and director of scientific 
programs. In 1983 he served as President of 
the American Microscopical Society; in 1990 
he was elected President of the World 
Association of Copepodologists. He is a 
Fellow of the American Academy of Arts and 
Sciences and a Research Associate at the 
Museum of Comparative Zoology of Harvard 
University. In 1982, William Jaspersohn, a 
popular writer of a series of photodocumentary 
books, selected Arthur among the many 
eminent marine biologists in Woods Hole to be 
the model of his new book “A Day in the Life 
of a Marine Biologist”. The book describes 
Arthur’s day, work in his office and laboratory, 
plus a field trip with students in his class in 
marine invertebrate zoology. It is a book very 
pleasantly read, about a kind and considerate 
gentleman who also is an excellent biologist 
and scholar. 
Arthur is well known among his associates, 
colleagues, and students as a kind and 
considerate gentleman. This courteous nature 
of Arthur is also revealed in his works on the 
symbiotic copepods. From time to time he 
would produce review articles or monographs 
for a group of copepods or a group of hosts 
with all of the reported copepod associates, in 
order to facilitate an easy way for the interested 
biologists to follow. Some notable examples of 
such works are in his reviews of the 
lichomolgid-complex, xarifid copepods, 
poecilostomatoids associated with soft corals, 
copepods of holothurians, and copepods 
associated with sea anemones. 
In his more than half-a-century affiliation with 
Boston University, the hardworking Arthur 
enthusiastically directed many of his students 
along the path of parasitology, copepodology, 
and marine invertebrate zoology that he had 
gingerly paved. Aside from being the teacher, 
mentor, and director of his students, Arthur 
served also the role of guardian to them. Every 
day at work, he would have in his office a tea 
time in the morning and a coffee break in the 
afternoon for his students to get together to 
relax, joke around, and talk about anything. 
Five of his former students followed his 
footsteps in copepodology, they are Roger F. 
Cressey, Masahiro Dojiri, Ju-shey Ho, John P. 
Murnane, and David C. Rosenfield. Arthur 
will be greatly missed by his friends and 
colleagues around the world in addition to his 
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