162 
The British Leeches 
Genus: Helobdella, R. Blanchard, 1896. 
Synonymy: 
Glossiphonia, Johnson, 1816 (partim). Clepsine, Savigny, 1817 (partim). 
Small Glossosiphonidae with one pair of eyes. Body generally 
without papillae. Complete somite composed of three rings. Crop with 
six pairs of simple lateral caeca, the last and longest pair reflected 
posteriorly. 
This genus has been established by Professor R. Blanchard (1896 a, 
p. 4) in order to admit Glossosiphonia stagnalis (Linnaeus, 1758) and a 
number of closely allied species which are found for the most part in 
South America. Johnson’s genus Glossosiphonia is thus reduced to a 
more homogenous group containing only six-eyed forms. 
Helobdella stagnalis, Linnaeus, 1758. 
Plate XIV, Figs. 13—17. Text Fig. 8 (p. 157). 
Synonymy and Literature: 
Rirudo bioculata, Bergman, 1757, pi. vi, figs. 9—11 ; 0. F. Muller, 1774, p. 41; 
0. F. Miiller, 1776, p. 220; Gmelin, 1788, p. 3096; Ure, 1793, p. 234; 
Schrank, 1804, p. 161; Braun, 1805, pp. 53—55, pi. vi, figs. 1—5 (coloured); 
Bose, 1802, p. 256 ; Stewart, 1817, p. 357 ; Carena, 1820, p. 302, pi. xii, fig. 21 
(coloured) ; Bruguitre, 1824, p. 132, pi. li, figs. 9—11. 
Hirudo stagnalis, Linnaeus, 1758, p. 649 ; Linnaeus, 1761, No. 2081 ; Weser, 1765, 
p. 44 ; Turton, 1806, p. 69 ; Turton, 1807, p. 129 ; Pennant, 1812, p. 71; Dalyell, 
1853, p. 36, pi. iv, figs. 1—16 (coloured). 
Hirudo pulligera, Daudin, 1800, p. 19, pi. i, figs. 1—3 ; de Blainville, 1827, p. 266. 
Hirudo circulars, Sowerby, 1806, p. 31, pi. lxxvi; Turton, 1807, p. 129; Pennant, 
1812, p. 72; Johnson, 1816, p. 27. 
Helluo ( Hirudo) biocxdata, Oken, 1815, p. 368. 
Glossiphonia perata, Johnson, 1816, p. 26. 
Glossopora punctata , Johnson, 1825, p. 50, pi. xvii, figs. 11—13. 
Erpobdella bioculata, de Blainville (in Lamarck), 1818, v, p. 296 ; de Blainville, 1827, 
p. 265. 
Clepsine bioculata, Savigny, 1822, p. 119; Carena, 1820; Moquin-Tandon, 1826, 
p. 102, pi. iv, fig. 2 ; Leydig, 1849, pi. iii, figs. 9—11 ; Diesing, 1850, p. 448 ; 
MetschnikofF, 1871, p. 505 (development) ; Whitman, 1878, p. 2 et seq. (develop¬ 
ment) ; Apathy, 1888a, p. 154 et seq., pi. viii, figs. 4 and 10 (head region); Apathy, 
1888 b, p. 790 et seq. (diagnosis); Leuckart, p. 610, fig. 260 b (proboscis); Oka, 
1894, p. 81 et seq. (sinus system, nephridia). 
Glossopora bioculata, Fleming, 1822, p. 604. 
Hirudo stagnorum, Derheims, 1825, pp. 10 and 20. 
Clepsine sowerbyi, Moquin-Tandon, 1826, p. 107 ; Diesing, 1850, p. 451. 
Hirudo ( Glossobdella ) pulligera, de Blainville, 1827, p. 266, pi. xxxvii, fig. 6. 
