150 
Haemogregarines 
after he had written his papers. Dr Sambon then goes on to adopt his 
usual method of bringing in everything he knows that may have even 
the remotest bearing on the subject, regardless of the fact that most of 
what he says has nothing to do with the question I raised. What I 
have said is quite clear, and is in no way connected with the multiplica¬ 
tion forms of other haemogregarines, yet Dr Sambon infers that I doubt 
whether other haemogregarines have such stages, and adds a list of 
names of observers, two of whom, as far as I know, have not described 
any of the multiplication forms of these parasites. I need hardly say 
that I have studied these stages of most of the haemogregarines. 
Now with regard to the vexed question of the sexual cycle of haemo¬ 
gregarines, Dr Sambon says under the heading “conjugation,” “ I can 
fully confirm Labbd’s observations, having witnessed it not only in vitro 
but also in blood taken from the gut of a tick fed on an haemogregarine- 
infected lizard”; nothing further is said about this parasite, which I 
presume is H. ehrlichi. Dr Sambon continues, “I have had the oppor¬ 
tunity of examining the process of accouplement so frequently in 
Haemogregarina seligmanni that I have no doubt whatever about it.” 
On the same page there are three figures which are not named, and 
which are not even referred to in the text, I can only presume therefore 
that they represent the sporonts of H. seligmanni in accouplement. 
They appear to have proceeded only a little way towards conjugation, 
and Dr Sambon himself says, “ I have not seen the nuclei of the con¬ 
jugating haemogregarines unite.” Yet he thinks this must take place in 
the gut of the invertebrate host, in this case I suppose in Porocephalus 
crotali. After having brought the reader breathlessly up to this point, 
eagerly expecting to have the mystery of the sexual cycle of these 
haemogregarines solved, Dr Sambon suddenly digresses to describe such 
uninteresting points as the structure and motility of the free sporonts. 
There is no proof that the parasites he figures are actually undergoing 
the process of conjugation, and I have certainly never seen any such 
process in snake haemogregarines, either in vitro, or in the alimentary 
tracts of ticks and linguatulids. Nothing more is said about the 
conjugation and further stages in the sporogony of “ H. seligmanni 
Dr Sambon however says in a footnote that they will be described in a 
future paper. Under the heading “sporogouy,” Dr Sambon merely 
re-describes the observations of Simond (1904), Durham (1902), 
Billet (1904), Brumpt (1904), Christophers (1905), Laveran and Negre 
(1905), and Prowazek (1908). With these conflicting statements and 
his limited observations on snake haemogregarines, Dr Sambon does 
