154 
Haemogregarines 
the herpetomonas stages of trypanosomes of vertebrates in the alimentary 
tracts of diptera, I hardly think he is in a position to criticise my work 
along these lines until he can prove that vertebrate trypanosomes 
have such stages; perhaps he may then be able to say I am wrong. 
As I have so far been unable to find any developmental stages of 
haemogregarines in invertebrates I can only tentatively accept Miller’s 
conclusions. It remains to be proved whether these blood parasites 
of mammals and reptiles will eventually be found to be transmitted in 
the extraordinary way suggested by Miller’s recent work. I am cer¬ 
tainly not so sanguine about it as Dr Sambon is. A great deal of work 
has yet to be done before it can be said for certain that the haemo- 
gregarine ookinete encysts and produces sporozoites in secondary cysts 
or sporebags. 
I will now give the last example of how Dr Sambon interprets other 
workers’ observations. In speaking of Leucocytozoon funambuli, he says, 
“In 1906, Captain Patton, in describing Haemogregarinci funambuli, 
a parasite of the five-striped palm squirrel {Funambulus pennantii), stated 
that he had seen free sporonts in the stomach of the squirrel’s louse, 
also a species of haematopinus.” If Dr Sambon will refer to my (1906) 
memoir on this parasite he will find that I have nowhere stated that I 
had seen the free sporonts of the parasite; Dr Sambon apparently 
knows more about it than I do. In the last Report of the Bacteriological 
Section of the King Institute I stated, “ I have again failed to find any 
extra-corporeal cycle in the lice found on Funambulus pennantii.” 
Dr Sambon either does not know of this Government publication or 
else he chooses deliberately to ignore it. How can he then in face 
of these facts say the free vermicules of L. funambuli are the sporonts 
of the parasite ? 
In a footnote to his paper, Dr Sambon expresses great surprise at 
Professor Nuttall permitting the use of the name Leucocytozoon in his 
Journal, and says, “ The name Leucocytozoon cannot be used for the 
haemogregarines of mammals whether they be parasites of leucocytes or 
not. It is the generic name of certain avian parasites discovered first by 
Danilewsky in 1884 in the blood of owls.” Dr Sambon will find my 
reason for retaining these parasites in this genus in my paper on 
“Mammalian Leucocytozoa” in the Report of the King Institute for 1907. 
With Laveran I prefer to place the avian parasites in the genus 
Haemamoeba ; the justification for this is fully supported by Mezincescu’s 
(1909) recent work. In conclusion I may say I do not intend wasting 
my time in further discussing with Dr Sambon the points I have 
raised, even though he should choose to reply to this paper. 
