E. Hindle and R. C. Lewis 
113 
beginning from the anterior extremity (Fig. 11), as usual in these 
flagellates. The two daughter parasites often remain attached for 
some little time before finally separating. 
Post-flagellate forms. The only parasite which appears to be a 
stage in the formation of these post-flagellate forms is that represented 
in Fig. 15. In this case the protoplasm is concentrated towards the 
posterior extremity and at one side there is a slight thickening of the 
ectoplasm suggesting the formation of a cyst wall. This parasite, however, 
differs from its neighbours in tbe presence of darkly staining gr anules 
and therefore may be a degenei’ating form. 
The fully formed cysts (Figs. 16, 17) are usually spherical bodies 
about 3 g in diameter, but in one case the dimensions of 3'5 /a by 6 /a were 
attained. In both cases these dimensions do not include the thickness 
of the cyst wall. 
Each cyst consists of a mass of densely staining cytoplasm containing 
the tropho- and kineto-nuclei; the end-bead is apparently absent. 
The cyst is surrounded by a cyst wall which in consequence of its non¬ 
staining properties appears as a hyaline layer. From a consideration of 
the life-cycle of other species of “ Grithidia,” presumably these are the 
forms which are excreted and serve to convey the infection to other 
hosts. 
We are in some doubt as to what generic name to apply to this 
species. There is no doubt that it falls under Patton’s definition of the 
genus Grithidia (1908), but as one of us has shown (Hindle, 1912), until 
the type species of Leptomonas, Kent, has been re-examined it is im¬ 
possible to apply any generic name to these flagellates! 
For the sake of convenience, however, we are placing it provisionally 
in the genus Grithidia {sensu Patton), and in view of its host, the 
specific name cleti is proposed. ‘^Grithidia'’ cleti is a flagellate 
occurring in the intestinal tract of Gletus varius Dali. It is not a 
stage in the life-cycle of any trypanosome occurring in the blood of 
vertebrates, for its host feeds entirely on the juices of plants. 
REFERENCES. 
Hindle, E. (1912). IVhat is the genus Leptomonas, Kent? Parasitology, v. 128. 
Patton, W. S. (1908). The life cycle of a species of Grithidia parasitic in the 
intestinal tract of Gerris fossarum Fabr. Arch. f. Protistenkunde, xii. 131-146. 
Parasitology v 
8 
