W. H. Leigh-Sharpe 
269 
curved claw with an accessory claw at its base ventrally and two smaller 
claws at the base on the side of the concavity. Near the end of the 
terminal joint there is also a rounded cushion of papillated spines, 
which, when the joints are brought together, would be nearly, but not 
quite, opposite the more distal cushion on the basal joint. The maxilli- 
pedes have lost their function of attaching organs, and are used only 
for pulling the head down to the skin of the host and holding it while 
the parasite feeds. 
The Mouth parts. (Fig. 5.) The mouth is tubular and forms a 
suctorial proboscis attached between the bases of the second antennae. 
It is situated at the summit of a cone, and is directed downwards and 
forwards against the skin of the host. There is ,an upper and a lower 
lip each of which is bordered by numerous setae of characteristic shape. 
The lips are loosely joined along their sides to form a tube by the split 
edges of the lower lip overlapping the edge of the upper on either 
side. In Fig. 5 the bps are shown separated. The under lip is the 
larger and more spoon-shaped so that its anterior end is curved like a 
horseshoe. 
The Alimentary canal, like that of many parasites, is degenerate 
and presents no special features. No glands appear to be present so 
that digestion is probably intra-cellular. The anus is not covered by 
plates or lamellae. I have always found the alimentary canal practically 
empty. Presumably the animal feeds on the soft tissues of the host 
which is not protected in the extra-cloacal aperture by the well-known 
dermal denticles. The pads of spines on the maxillipedes are well 
adapted for rasping any parts of the host, though these appendages 
are probably used for closer attachment of the head during feeding. 
The Reproductive system consists of a pair of ovaries on either 
side of the alimentary canal. These are seen full of eggs. Besides 
which, as is usual in the Copepoda, the eggs descend into ovisacs, or 
egg-sacs (Fig. 2), which are situated dorsal to the genital processes and 
are long and slender, of equal length with or at least five-sevenths the 
length of the trunk. The eggs are large and flattened to the form of 
oblate spheroids. There are in the ovisacs usually four to six rows of 
22 or 23 eggs each 1 . 
The oviducts leave the ovaries about half-way down on their outer 
borders. The Cement glands are large and crescentic and are attached 
1 This appears a very average condition, and agrees with Scott’s figure of L. galei, 
while Wilson gives four to six rows of about 150 eggs in each string in L. galei, at the same 
time stating that the ovisacs are as long as the body. 
