L. Harrison and T. H. Johnston 
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Sub-family BOOPINAE n. 
Characters of sub-family. Boopidae with four-jointed palps, and 
long, two-jointed tarsus. Antennary fossa always bounded above and 
below by chitinous plate-like expansions of the sides of the head, both 
deeply cleft about the middle of their length. The eye is situated in the 
dorsal plate, just behind the cleft. Mesothoracic prominences always 
with stout spines. Three pairs of special sensory organs upon the first 
three abdominal segments. 
Three genera are known, Boopia Piaget, Heterodoxus Le Souef and 
Bullen, and Paraheterodoxus here described as new. The three may 
be easily distinguished by means of the following key. 
Fig. 4. First three abdominal segments of Heterodoxus longitarsus, to show special 
sensory hairs (H 1 , H 2 , H 3 ). (L. H. del.) 
Key to the Genera of Boopinae. 
A. Without dagger-like spines beneath head Boopia. 
AA. With dagger-like spines beneath head ... B. 
B. Spines rising from base of palps ... ... Heterodoxus. 
BB. Spines rising from throat beneath occiput Paraheterodoxus. 
Genus Boopia Piaget. 
The genus Boopia has been well enough characterised by Piaget 
(1880, p. 599), but we would emphasise the special features which 
distinguish this genus, which comprises more species than any other of 
the family, from the bird-frequenting forms. These features are 
possessed in common by the remaining genera of the family, but Boopia 
looks distinct to the ordinary systematise while Heterodoxus has been 
