PAN-PACIFIC ENTOMOLOGIST 
Vol. 55, No. 1, pp. 10-20 
THE LARVAL STAGE OF HYDROPSYCHE SEPARATA BANKS 
(TRICHOPTERA: HYDROPSYCHIDAE) 
Douglas Smith 
Dept, of Biol., Univ. of Saskatchewan, Saskatoon, Canada S7N 0W0 
There is some confusion concerning the name used for the species which 
Banks (1936) described under the name Hydropsyche separata Banks. 
Banks (1936) described the male of H. separata and Denning (1943) de¬ 
scribed the female, illustrated the female genitalia and provided a more 
complete drawing of the male genitalia of this species. Ross and Spencer 
(1952) suggested H. separata described from North America was a synonym 
of the European species Hydropsyche guttata Pictet. Ross (1965), in a pri¬ 
marily zoogeographical paper, stated that H. guttata and H. separata were 
distinct species. Personal communications with other Trichoptera taxono¬ 
mists and the published literature (e.g., Baumann and Winget, 1975), how¬ 
ever, indicate that the name H. guttata is still used to refer to the species 
H. separata. 
Discussion 
Dr. Hans Malicky (from the Biologische Station Lunz, Lunz, Austria) 
kindly sent me male specimens of H. guttata and I sent him males of H. 
separata. After comparing these two species we concluded that they are 
distinctly different (even the number of tibial spurs differ in the males). This 
conclusion supports Ross’s (1965) view that the species H. guttata and H. 
separata are separate species and that the name H. guttata should not be 
used to refer to the species H. separata which Banks described in 1936. 
Ross (1965) suggested that H. separata and H. guttata are daughter 
species of a common ancestor but Dr. Malicky (pers. comm., 1977) found 
that, based on male specimens, H. separata is more closely related to a 
European species which he had recently described (Malicky, 1977), Hydro¬ 
psyche bulgaromanorum Malicky, than to H. guttata. 
Dr. A. P. Nimmo (oral comm., 1978) notes that the species Hydropsyche 
corbeti Nimmo which he described (Nimmo, 1966) may be a synonym of 
Hydropsyche separata. The drawing of the male genitalia of H. separata 
in Denning (1943) and the male genitalia of specimens of H. separata which 
I examined are extremely similar to the drawings of the male genitalia of H. 
corbeti presented by Nimmo (1966). Since I have not, as yet, examined the 
type of H. corbeti, I can only state that my observations on the close 
