10 
“TERRA NOVA'’ EXPEDITION. 
way distinctive are the “ perfectly-segmented ” body and the “ nine-jointed ” palps.* 
The first character is shared by two species of Colossendeis, C. articulata and C. dofieini 
of Loman (1908, p. 22, and 1911, p. 4). As regards the second character, N. germanica 
agrees in this, but, apparently, in no other respect, with the species described below as 
C. ivilsoni. If the genus were to be retained it would be hard to decide whether it 
should include C. articulata and C. dofieini on the one hand or C. wilsoni on the other; 
it could not embrace all three. 
A large number of species of Colossendeis have been described, most of them from 
a very small number of specimens. They are distinguished mainly by proportional 
differences of measurement, and there is reason to believe that some of them would not 
survive a critical revision based on adequate collections. In the absence of such a 
revision it is necessary, before venturing to describe any additional species, to attempt 
to reduce to some sort of order those already known. Bouvier has made a beginning 
by dividing the species into two groups according to the relative lengths of the distal 
segments of the legs. In the first or “ longitarsal ” group the carpus, propodus, and 
claw together measure at least three-quarters of the length of the second tibia ; in the 
“ brevitarsal ” group the proportion is always very much less. Proceeding on these 
lines, the following key may be offered for the “ longitarsal ” group, which includes all 
the Antarctic species. 
Key to the “ Longitarsal ” species of Colossendeis. 
a. Sixth segment of palp more than three times as long as thick. Proboscis 
distinctly longer than trunk. 
a'. Trunk segmented ......... 
V. Trunk not segmented. 
a". Lateral processes in contact. 
a”'. Seventh segment of palp longer than eighth. Eyes absent 
b "'. Seventh segment of palp shorter than eighth. Eyes present 
b". Lateral processes separated. 
a'". Seventh segment of palp equal to eighth .... 
b'". Seventh segment of palp distinctly shorter than eighth. 
a'"'. Eyes absent. 
Proboscis dilated distal ly .... 
Proboscis dilated in middle .... 
C. articulata, Loman 
C. prroboscidea (Sabine) 
C. scotti, sp. n. 
C. australis, Hodgson 
( C. media, Hoek f 
(C. brevipes, Hoek 
C. orcadensis, Hodgson 
f (1. angusta, G. O. Sars f 
(C. gracilis, Hoek 
* I learn from Mr. Hodgson that he does not accept the view of Bouvier (1913, p. 37), according to 
which only nine segments are counted in the palp of normal species of Colossendeis. Bouvier, no doubt 
rightly, excludes from the enumeration the “saillie basilaire ” of the palps, which is generally counted as 
a segment, although it is precisely similar to the process (never reckoned as a segment) that lies alongside 
it and carries the oviger. Loman is said by Bouvier to have been the first to call attention to this point, 
but I cannot discover the passage in which he did so. 
f C. media and C. brevipes were described by Hoek as doubtfully distinct from C. gracilis, which 
again is identified by Mobius with C. angusta. Meinert’s observation (1899, p. 59) as to the differences in 
form of the palpal segments in immature and fully adult specimens of the last-named species deserves to 
be noted as having possibly a wider application. 
