Isler and Whitney • SPECIES LIMITS IN AN ANTBIRD COMPLEX 
13 
Vriesendorp, C., T. S. Schulenberg, W. S. Alverson, D. 
K. Moscovits, and J.-I. Rojas Moscoso. 2006. Peru: 
Sierra del Divisor. Rapid Biological Inventory Report 
17. The Field Museum, Chicago, Illinois, USA. 
Willis, E. O. 1982. The behavior of Scale-backed Antbirds. 
Wilson Bulletin 94:447—462. 
Zimmer, J. T. 1934. Studies of Peruvian birds. XII. Notes 
on Hylophylax , Myrmothera, and Grallaria. American 
Museum Novitates 703:1-21. 
Zimmer, K. J. and M. L. Isler. 2003. Family Thamnophi- 
lidae (typical antbirds). Pages 448-681 in Handbook 
of the birds of the world. Volume 8. Broadbills to 
tapaculos (J. del Hoyo, A. Elliott, and D. A. Christie, 
Editors). Lynx Edicions, Barcelona, Spain. 
APPENDIX 
Recordings Examined. —The following list 
identifies recordings used in the study by taxon, 
country, state or department, recording location, 
and recordist. Numbers following the recordist 
name identify the number of cuts per recordist per 
location. Acronyms for recording archives: BSA 
= Banco de Sonidos Animates, Instituto de 
Investigation de Recursos Biologicos Alexander 
von Humboldt, Bogota. FSM = Florida State 
Museum Sound Archive. ML = Macaulay 
Library, Cornell Laboratory of Ornithology, 
Ithaca; ISL = recordings not yet archived in an 
institutional collection but that have been copied 
into the inventory maintained by Morton and 
Phyllis Isler. Many of these unarchived recordings 
either are in the process of being archived or will 
eventually be archived by the recordists. Nomen¬ 
clature reflects recommended taxonomic position. 
Willisornis p. poecilinotus : (53 recordings; 24 
locations). Brazil: Amapa: Porto Grande (Zimmer 
3 ISL); Amazonas: Rio Apuau (Cohn-Haft 1 ISL), 
60-90 km N Manaus (Bierregaard 1 ML, Stouffer 
1 ML, Whitney 1 ISL, Whittaker 1 ISL), Reserva 
Ducke (Whitney 1 ISL). Guyana: Acarai Moun¬ 
tains (Robbins 1 ML), Baramita (Brumfield 1 
ISL), Iwokrama Forest Reserve (Whitney 5 ISL), 
Kaieteur Fall (Milensky 1 ISL), Kako River 
(Robbins 2 ML), Kopinang (O'Shea 1 ML), 
Kuyuwini River (Finch 2 ML), Marshall Falls 
(Finch 1 ML), Nappi Village (Parker 1 ML), Sipu 
River (Milensky 1 ISL), Waruma River (O’Shea 1 
ML). Suriname: Brownsberg Nature Reserve 
(Davis 2 ML, Whitney 2 ISL), Kraka-Zenderij 
Road (Whitney 1 ISL), Raleigh Vallen (Whitney 1 
ISL), Voltzberg (Davis 1 ML, M. Isler 1 ML, 
Whitney 1 ISL). Venezuela: Amazonas: Jungla- 
ven Camp (Zimmer 1 ISL); Bolivar: El Palmar 
(Parker 2 ML, Schwartz 3 ML), La Escalera 
(Behrstock 1 ISL, M. Isler 1 ML, Macaulay 1 ML, 
Schwartz 1 ML, Whitney 1 ISL, Zimmer 2 ISL), 
20-30 km SE Maripa (Stejskal 1 ISL, Whitney 2 
ISL), Sierra de Lema (Behrstock 1 ISL, Zimmer 1 
ISL). 
Willisornis p. duidae : (29 recordings; 9 loca¬ 
tions). Brazil: Amazonas: P. N. do Jaii (Pacheco 1 
ISL, Whitney 1 ISL, Whittaker 1 ISL), Manaca- 
puru (Whitney 1 ISL), Sao Gabriel da Cachoeira, 
island in Rio Negro (Whitney 2 ISL), Sao Gabriel 
da Cachoeira. L bank Rio Negro (Whittaker 1 ISL, 
Whitney 3 ISL, Zimmer 2 ISL), Sao Gabriel da 
Cachoeira. R bank Rio Negro (Cohn-Haft 1 ISL, 
Whitney 1 ISL, Zimmer 1 ISL). Colombia: P. N. N. 
Chiribiquete (M. Alvarez 10 BSA), Vaupes: Mitu 
(Hilty 2 ISL). Venezuela: Amazonas: Frente Isla 
Cigarron (Schwartz 1 ML), Picua (Coons 1 ISL). 
Willisornis p. duidae or lepidonotus: (4 record¬ 
ings; 1 location). Colombia: Amazonas: Parque 
Nacional Natural Amaca-yacu (P. Isler 1 ISL, 
Whitney 3 ISL). 
Willisornis p. lepidonotus below 800 m: (37 
recordings; 18 locations). Ecuador: Morona-San- 
tiago: Miazal (Whitney 1 ISL), Santiago (Robbins 
1 ISL); Napo: La Selva Lodge (Behrstock 1 ISL, 
Coopmans 2 ML, Donahue 1 ML, Wolf 2 ISL), 
km 37 Maxus Road (Krabbe 3 ISL), Tiputini 
Biodiversity Station (Behrstock 1 ISL, Zimmer 2 
ISL); Pastaza: Kapawi Lodge (Whitney 1 ISL, 
Wolf 1 ISL); Sucumbtos: Cuyubeno (Whitney 1 
ML). Peru: Loreto: Colonia Angamos (Lane 1 
ISL), 79 km WNW Contamana (Lane 3 ISL), El 
Dorado (Whitney 3 ISL), El Tigre (J, Alvarez 3 
ISL), Explorama Lodge (Whitney 1 ISL), Intuto 
(Whitney 1 ISL), Quebrada Oran (Whitney 2 
ISL), Quebrada Sucusari (P. Isler 1 ML), 
Sachacocha (J. Alvarez 1 ISL), Yanamono 
(Budney 2 ML, Whitney 1 ISL); San Martin: 
Tarapoto- Yurimaguas Road (Lane 1 ISL). 
Willisornis p. lepidonotus above 800 m: (16 
recordings; 8 locations). Ecuador: Napo: 15 to 
80 km W of Loreto by road (J. Rowlett 1 ISL, R. 
Rowlett 1 ISL, Whitney 2 ISL, 1 ML, Wolf 1 
ISL), Volcan Sumaco (R. Rowlett 1 ISL); 
Zamora-Chinchipe: Parque Nacional Podacaipus 
(Wolf 1 ISL). Peru: Cajamarca: Cordillera del 
Condor, (Schulenberg 1 ISL), Puesta Vigilancia 
(Schulenberg 1 ML); Loreto: 77 km WNW 
Contamana (Lane 3 ISL), -90 km SE Juanjui 
(Lane 2 ISL); San Martin: Jirillo, 15 km NE 
(Schulenberg 1 ML). 
Willisornis p. gutturalis : (12 recordings; 3 
locations). Brazil: Amazonas: Benjamin Constant 
(Whitney 4 ISL), R. N. Palmarf (Whitney 5 ISL, 
