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THE WILSON ORNITHOLOGICAL SOCIETY 
FOUNDED 3 DECEMBER 1888 
Named after ALEXANDER WILSON, the first American ornithologist. 
I 
President —E. Dale Kennedy, Biology Department, Albion College, Albion, MI 49224, USA; e-mail: 
dkennedy@albion.edu 
First Vice-President —Robert C. Beason, P. O. Box 737, Sandusky, OH 44871, USA; e-mail: 
Robert. C .Beason@gmail .com 
Second Vice-President —Robert L. Curry, Department of Biology, Villanova University, 800 Lancaster 
Avenue, Villanova, PA 19085, USA; e-mail: robert.curry@villanova.edu 
Editor —Clait E. Braun, 5572 North Ventana Vista Road, Tucson, AZ 85750, USA; e-mail: TWILSONJO@ 
comcast.net 
Secretary —John A. Smallwood, Department of Biology and Molecular Biology, Montclair State University, 
Montclair, NJ 07043, USA; e-mail: smallwoodj@montclair.edu 
Treasurer —MelindaM. Clark, 52684 Highland Drive, South Bend, IN46635,USA; e-mail: MClark@tcservices.biz 
Elected Council Members —Jameson F. Chace, Sara R. Morris, and Margaret A. Voss (terms expire 
2011); Mary Bomberger Brown, Mary Garvin, and Mark S. Woodrey (terms expire 2012); Mark 
Deutschlander, Paul G. Rodewald, and Rebecca J. Safran (terms expire 2013). 
Living Past-Presidents —Pershing B. Hofslund, Douglas A. James, Jerome A. Jackson, Clait E. Braun, 
Mary H. Clench, Richard C. Banks, Richard N. Conner, Keith L. Bildstein, Edward H. Burtt Jr., John 
C. Kricher, William E. Davis Jr., Charles R. Blem, Doris J. Watt, and James D. Rising. 
Membership dues per calendar year are: Regular, $40.00; Student, $20.00; Family, $50.00; Sustaining, 
$100.00; Life memberships, $1,000.00 (payable in four installments). 
The Wilson Journal of Ornithology is sent to all members not in arrears for dues. 
THE WILSON JOURNAL OF ORNITHOLOGY 
(formerly The Wilson Bulletin ) 
THE WILSON JOURNAL OF ORNITHOLOGY (ISSN 1559-4491) is published quarterly in March, June, 
September, and December by the Wilson Ornithological Society, 810 .East 10th Street, Lawrence, KS 66044-8897. The 
subscription price, both in the United States and elsewhere, is $40.00 per year. Periodicals postage paid at Lawrence KS. 
POSTMASTER: Send address changes to OSNA, 5400 Bosque Boulevard, Suite 680, Waco, TX 76710. 
All articles and communications for publication should be addressed to the Editor. Exchanges should be addressed 
to The Josselyn Van Tyne Memorial Library, Museum of Zoology, Ann Arbor, MI 48109, USA. 
Subscriptions, changes of address, and claims for undelivered copies should be sent to OSNA sunn 
Boulevard. Suite 680. Waco, TX 76710, USA. Phone: (254) 399-9636; e-mail: business@osnabirds org Back issues or 
single copies are available for $12.00 each. Most back issues of the journal are available and may be ordered from OSNA 
Special prices will be quoted for quantity orders. All issues of the journal published before 1117, 0 2 
web site a, the University of New Mexico library (http://elibrary.u,Ledu/L^ 
reproductions of all papers (including illustrations) are available as either PDF or DjVu files ^ hable > and fiiH-text 
© Copyright 201 I by the Wilson Ornithological Society 
Printed by Allen Press Inc., Lawrence, KS 66044, USA 
COVER: Wilson’s Snipe (Gallinago delicaia). Illustration by Scott Rashid 
© This paper meets the requirements of ANSI/NISO Z39.48-1992 
(Permanence of Paper). 
