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THE WILSON JOURNAL OF ORNITHOLOGY • Vol 123, No. 4. December 2011 
The BUK population is classified as intern 
diate between the KAR-KAZ complex and AL 
This agrees with both the refugia hypothesis 
later partial reconnection and the hypothesis 
isolation by distance only. The Great Re 
Warbler {Acrocephalus antndinaceus) populate 
in the Eurasian breeding range of the species h 
two dades .hat have colonized from two separate 
n? 9 nno ,n S ir0pe Und the Middlc Easl (Hansson 
aL 20°S,. The forest lypes of KAR and KAZ we 
black pine while the o.her populations occupit 
affected the"'' T f ° rrat W* 1 •!*> hm 
d the morphometries independently ( 
isolation due to long distances or refugia discon¬ 
nection between locations ALA and the KAR- 
KAZ complex (min = 440 km: Fig. 1). 
Anatolian refugia are known to have affected 
the evolution of structured populations in several 
groups including grasshoppers (Ciplak 2008 k 
small mammals (Castigfia et al. 2009). and a 
passerine bird (Hansson et al. 2008 ). For example- 
the Antrastes genus of Orthoptera arose from an 
ancestral stock during the Pliocene era in 
northwestern Anatolia whose range later expand 
ed and contracted under the effects of the ice ages 
(glacial and interglacial periods, respectively! 
