36 
Card. Bull. Singapore 69(1) 2017 
Notes. Hoya australis R.Br. ex J.Traill is a widely variable species thought to occur 
in Australia, Papuasia and Melanesia (Hill, 1988; Forster & Liddle, 1991; Liddle & 
Forster, 2008). It is currently classified into seven subspecies based on vegetative 
morphology and distribution (Liddle & Forster 2008; Kloppenburg, 2012). The 
discovery of Hoya australis in Sabah (based on Jamirus, J. in Lamb , A. AL2500/2014, 
SAN) considerably extends its distribution area. Preliminary molecular analysis 
confirms that the Bornean collection belongs to Hoya australis but it is inconclusive 
to its sub-specific ranking (Rodda, unpublished). This is the reason why synonymies 
are not indicated here. For the most updated treatment of subspecies and synonymies 
of Hoya australis see Liddle & Forster (2008). 
The specimen here indicated as lectotype of Hoya australis has long been 
considered a holotype (Hill, 1988; Smith, 1988; Forster & Liddle, 1991; Liddle & 
Forster, 2008). The specimen is not a holotype as the protologue of Hoya australis is 
not more specific than that the original material is from Australia in Brown’s herbarium. 
It is therefore impossible to establish whether it was based on a single specimen only. 
Hill (1988), in mentioning Banks , J. & Solander, D. s.n. (BM) as holotype of H. 
australis , is here considered as an effective lectotypification following ICN Art. 9.9 
(McNeill et al., 2012). 
Hoya bakoensis Rodda, Phytokeys 53: 87 (2015) [21 July 2015]. - TYPE: Malaysia, 
Sarawak, Bako National Park, 20 March 2015, Rodda, M. MR1042b (holotype SING; 
isotypes SAR, KEP). 
Hoya beccarii Rodda & Simonsson, Webbia 68: 13 (2013). - TYPE: Malaysia, 
Sarawak, Matang, July 1866, Beccari, O. 1866 ( collezioni Beccari 6526a) (holotype 
FI; isotype FI). 
Hoya benchaii Gavrus, A.L.Lamb, Emoi & Gokusing, Sandakania 19: 18 (2014). - 
TYPE: Originally from Malaysia, Sabah, east of Ranau, Telupid area, cultivated at 
Kipandi Park, vouchered on 8 May 2011 as Chai, B. in Lamb, A. 2132/2011 (holotype 
SAN; isotype SNP n.v.). 
Hoya callistophylla T.Green, Fraterna 13(4): 2 (2000). - TYPE: Originally from 
Malaysia, Sabah, Nabawan, cultivated in USA, Hawaii, Oahu, Ka‘a‘awa, vouchered 
on unknown date as Green , T. 201 (holotype BISH [BISH1014775]). 
Hoya caudata Hook.f., FI. Brit. India 4(10): 60 (1883); Rintz, Malayan Nat. J. 30: 
484 (1978). - TYPE: Peninsular Malaysia, Malacca, Maingay, A.C. 1956 (Kew distr. 
no. 1128) (lectotype K [K000895134], designated by Rintz (1978), first step, and 
designated here, second step; isolectotypes K [K000895133], L [L0004315]). 
Hoya crassifolia Ridl., J. Straits Branch Roy. Asiat. Soc. 61: 30 (1912), nom. illeg., 
non Hoya crassifolia (J.Jacq.) Haw. - Hoya caudata Hook.f. var. crassifolia Ridl., FI. 
Malay Penins. 3: 395 (1923). - TYPE: Malaysia, Perak, Tapah, on the trees on the 
Temoh Road, November 1908, Ridley, H.N. 14059 (lectotype SING [SING0059473], 
designated here; isolectotype K [K001096330]). 
