Names and types of Bornean Hoya 
37 
Notes. Hooker indicated the type of Hoya caudata Hook.f. as Maingay 1128. This 
is the K distribution number of the Maingay sheets that corresponds to Maingay 
1956. The indication by Rintz (1978) of Maingay 1128 (K) as ‘type’ is an effective 
lectotypification under Art. 9.9 of the ICN (McNeill et al., 2012). However in K there 
are two sheets labelled Maingay (.Kew distr. no. 1128 ) and, therefore, a second-step 
lectotypification is necessary. The duplicate with barcode K000895134 is a well- 
preserved fertile specimen that also bears a pencilled sketch and is here selected as 
lectotype of H. caudata. 
Two duplicates have been found of the type of Hoya crassifolia Ridl. The SING 
sheet [SING0059473] is a well-preserved fertile specimen and is here selected as 
lectotype. 
Hoya chewiorum A.L.Lamb, Gavras, Emoi & Gokusing, Sandakania 19: 25 (2014). 
-TYPE: Originally from Malaysia, Sabah, Central Crocker Range, Sg. Kopongian on 
the upper reaches of the Moyong River, cultivated at Kipandi Park, vouchered on 30 
April 2009 as Gokusing, L. andLombika, J. in Lamb, A. AL1769/2009 (holotype SAN; 
isotype SNP n.v.). 
Hoya clemensiorum T.Green, Fraterna 14(3): 12 (2001). - TYPE: Originally from 
Malaysia, Sabah, Poring, Rafflesia Centre, 16 September 1991, cultivated in USA, 
Oahu, Ka‘a‘awa, Hawaii, reputedly vouchered on unknown date as Green, T. 91032 
(holotype BISH, missing); Malaysia, Sabah, Mt. Kinabalu, Dallas, 8 Sepember 1931, 
Clemens, J. & M.S. 26358 (neotype K, designated here; isoneotypes BM, BO, L). 
Notes. The holotype of Hoya clemensiorum T.Green, Green 91032 dated 16 
September 1991, cannot be found at BISH. In BISH there is instead a sterile specimen 
with the same collector and number dated 09 June 2008 labelled as neotype of H. 
clemensiorum , obtained in cultivation from a plant originally wild collected in Sabah 
on 16 September 1991.1 have been unable to find the publication of a neotypification 
and I therefore select Clemens 26358 (K) as the neotype of H. clemensiorum as it 
matches the original description, is fertile, and is among one of the best preserved 
specimens of H. clemensiorum collected by the Clemenses. 
Hoya coriacea Blume, Bijdr. FI. Ned. Ind. 16: 1063 (1826); Rintz, Malayan Nat. J. 30: 
495 (1978). -TYPE: Indonesia, Java, Salak [In fruticetis adpedemmontis Salak’], s.d., 
s.coll. (lectotype L [sheet number 898.168-117], designated here; possible isolectotype 
P [P00639838]). 
Hoya brunoniana Wight, Contr. Bot. India 37 (1834). - TYPE: Penang?, Wall Asclep. 
no 37[= Wall. Cat. 8163] (holotype K; isotypes K [K001129108], E [E00179575]). 
Hoya fraterna Blume, Rumphia 4: 32 (1849) [late October 1849]; Blume, Mus. Bot. 
1(3): 44 (1849) [November 1849], - TYPE: Indonesia, Java, West Java, Kuripan [In 
calcareis Kuripan Javae occidentalis] s.d., s.coll. (lectotype L [L0004318], designated 
here) (Synonymy established by Backer & Bakhuizen van den Br ink , 1965). 
