38 
Card. Bull. Singapore 69(1) 2017 
Hoy a occlusaRidl., J. Straits Branch Roy. Asiat. Soc. 61: 31 (1912). - TYPE: Malaysia, 
Selangor, top of the Batu Caves, December 1890, Ridley, H.N. s.n. (lectotype SING 
[SING0059478], designated here). 
Hoya angustisepala Elmer ex C.M.Burton, Hoyan 8(4) pt 2: 6 (1987). - Hoya 
mindanaensis Elmer, Leafl. Philipp. Bot. 10: 3585 (1938), nom. inval. (missing Latin 
description). - TYPE: Philippines, Davao, Mindanao, Todaya, Mt. Apo, Sibulan River, 
3000 ft., June 1909, Elmer, A.D.E. 10829 (lectotype B [B100277215], designated here; 
isolectotypes BISH [BISH1000876], BM [BM000945404], BO, E [E00288757], K 
[K000911125], L [L0004313], L(U) [U1102604],NY [NY00318664], US [00170056]). 
Notes. Blume did not directly indicate any specimen for Hoya coriacea Blume but 
mentioned the provenance ‘ In fruticetis ad pedem montis Salak/. I have located one 
specimen in L bearing an autograph label by Blume ‘ Hoya coriacea; Salak’ which is, 
therefore, suitable for designation as the lectotype. A specimen sent by Blume to P in 
1836 [P00639838] which is labelled ‘In fruticetis ad pedem montis Salak ’ is a perfect 
match to Blume’s description. However there is no autograph label and therefore the L 
specimen is preferred as lectotype for H. coriacea. 
Duplicates of Wall. Asclep. 37[= Wall. Cat. 8163] have been found at E and K. 
The K duplicate is the only one bearing a Herbarium Robert Wight Proper (HRWP) 
label, it is annotated ‘37 H. brunoniana ’ has a sketch of the flower in Wight’s hand and 
is therefore one of Wight’s specimens on which he based his descriptions. Linn Soc 
Mss SP1284 indicates that only 1 sheet of Wall. Asclep. 37 was given to Robert Wight 
for his personal working herbarium, therefore the K sheet can be considered a holotype. 
The name Hoya fraterna was first mentioned in volume four of Rumphia (Blume, 
1849: 32) [late October 1849] where Blume compared it to the similar species Hoya 
coriacea (fraterna = brotherly, closely associated (Steam, 2008)). The description is 
expanded in Blume (1849: 44) [November 1849] (Stafleu & Cowan, 1976). Blume 
did not refer to any specimens but mentioned the provenance as ‘In calcareis Kuripan 
Javae occidentals I have located one specimen in L bearing an autograph label in 
Blume’s hand ‘Hoyafraterna Bl; in calcareis Kuripan’ that is to be considered original 
material and is, therefore, selected as lectotype. 
One duplicate has been found of the type of Hoya occlusa Ridl. at SING 
[SING0059478], The specimen is well-preserved and fertile and is here selected as 
lectotype. 
Hoya coronaria Blume, Bijdr. FI. Ned. Ind. 16: 1063 (1826); Rintz, Malayan Nat. 
J. 30: 505 (1978). - Eriostemma coronaria (Blume) Kloppenb. & Gilding, Fraterna 
14(2): 1 (2001). - TYPE: Indonesia, Java, s.d., s.coll. ‘124/6’ (lectotype L, [sheet 
no. 898.168-121], designated by Kleijn & Van Donkelaar (2001: 469); isolectotype L 
[sheet no. 898.168-128]. 
Eriostemma obtusifolioides Gilding & T.Green, Fraterna 22(3): 5 (2009). - TYPE: 
Originally from Malaysia, Sabah, Apin Apin, cultivated in USA, Hawaii, Oahu, 
Ka‘a‘awa, Green Plant Research, vouchered in 2007 as Green, T. s.n. (holotype BISH 
[BISH1072387]). 
