Names and types of Bornean Hoya 
41 
Hoyapersicinicoronaria Shao Y.He & P.T.Ei, Novon 19: 475 (2009). - TYPE: China. 
Hainan Prov., Bawangling, on a tree in montane tropical rainforest, 19°06'N 109°06'E, 
214 m, 8 August. 2007, He, S.Y. & Lin, J.Y. 0708086 (holotype CANT, n.v., photo at 
MO not found). 
Notes. Blume (1826) indicated in the protologue of Hoya diversifolia a reference to 
Rumphius’ Herbarium Amboinense 5 Tab. 175 Fig. 2 and no direct or indirect reference 
to a specimen. Rumphius’s plate does not have a detailed illustration of the flowers and 
does not entirely match Blume’s description as the leaves, described as ‘aveniis’ by 
Blume have evident venation in the plate. However Blume’s description may be based 
on live plants, as fresh leaves do not have distinct venation while veins are sometimes 
visible in dry specimens. No specimen from Blume’s time annotated as collected in 
‘Kuripan’ or ‘circa Bataviam’ has been found. Herbarium Amboinense 5 Tab. 175 Fig. 
2 is therefore the only material available for lectotypification. In L there are numerous 
specimens identified as Hoya diversifolia with annotations in Blume’s hand. Among 
these there is one labelled Hoya heterophylla in Blume’s hand, a nomen nudum, 
and with a very similar etymology as Hoya diversifolia. Sheet number 989168-147 
is a well-preserved fertile specimen and is therefore designated as epitype for Hoya 
diversifolia. Specimen [P00639841] in Pis labelled both Hoya heterophylla and Hoya 
diversifolia Bl. Bijdr’ in Blume’s (or Decaisne?) hand, the locality is Java, ‘m.fisit] 
Blume 1836’ and may be original material, however it is impossible to verily if it was 
collected prior to 1826. 
Hoya crassipes Turcz. was based on Zollinger 2581. The KW duplicate 
[KW001000511] is a well-preserved fertile sheet and is designated here as lectotype. 
Hoya orbiculata Wall, ex Wight & Arn. was based on Wall. Asclep. 32 [= Wall. 
Cat. 8151]. A specimen belonging to what used to be Wight’s personal herbarium and 
bearing a sketch of the flower in Wight’s hand is present at K and is the most suitable 
lectotype for Hoya orbiculata (Noltie 2005: 134). The plate of Sussuela esculenta in 
Rumphius (1747: t. 175.28) is a syntype of Hoya orbiculata. 
Hoya esculenta Tsiang is an illegitimate name because it is clearly indicated as 
a synonym of the earlier Hoya diversifolia Blume and Hoya orbiculata Wall, ex Wight 
& Arn. and is therefore superfluous and illegitimate following ICN Art. 52.1 (McNeill 
et al., 2012). It is lectotypified by Herb. Amb. 5: 467, t. 175.28. 
The type of Hoya liangii Tsiang was indicated as Liang 26867. The IBSC 
duplicate [IBSC0005685] is a well-preserved fertile sheet and is designated here as 
lectotype. The syntype How, C.F. 73935 is also available at IBSC [IBSC0005687]). 
Hoya elliptica Hoolc.f., FI. Brit. India 4: 58 (1883); Rintz, Malayan Nat. J. 30: 505 
(1978). - TYPE: Malaysia, Malacca, 1 April 1868, Maingay, A.C. 3286, Kew Distrib. 
1137 (lectotype K [K000895126], designated here; isolectotype K [K000895127]). 
Notes. Hooker described Hoya elliptica citing specimen Kew Distrib. 1137. Two 
sheets are preserved at K labelled Kew Distrib. 1137 as well as bearing the original 
collection number Maingay 3286. The duplicate here selected as lectotype is well 
preserved, fertile and bears a dissection and a sketch of the flowers by Gamble as an 
aid to identification. 
