Names and types of Bornean Hoya 
49 
the first time in August 1838’. In Lindley’s herbarium at CGE there is a sheet labeled 
‘Hoya coriacea Loddiges Aug. 1830’. Excluding the date that does not match, this 
sheet matches the protologue and being the only original material available is here 
selected as lectotype for Cyrtoceras lindleyanum. Miquel (1857: 515) cited Cyrtoceras 
lindleyanum Miq. with himself as the author. This, however, is likely a typographical 
error as he clearly also included Cyrtoceras lindleyanum Decne. in the citation. 
Cyrtoceras floribundum is clearly indicated as a synonym of the earlier 
Cyrtoceras reflexum Benn. and is therefore superfluous and illegitimate following ICN 
Art. 52.1 (McNeill et al., 2012). The name was based, as per Hoya coriacea Lindl., 
on a live plant originally collected by Cuming in the Philippines and cultivated at 
Loddiges Nursery (UK). No specimens have been found with such provenance. The 
only material available for the selection of a lectotype is therefore the plate included 
together with the species description. 
No specimens are mentioned for Centrostemma micranthum Blume. The 
illustration published concurrently with the description is therefore selected as the 
lectotype. 
No specimens are mentioned for Centrostemma elegans Blume. There is, 
however, a specimen in L identified as Centrostemma elegans in Blume’s handwriting, 
also labelled Cyrtoceras elegans (Blume) Miq in Miquel’s hand and Hoya elegans 
(Blume) Boerl. in Boerlage’s hand, that is a suitable lectotype. 
No specimen was mentioned in the description of Centrostemma laurifolium, 
only ‘In sylvis montanis Javae occidentalis.’ A specimen bearing a label in Blume’s 
hand from Salak is here selected as lectotype. 
Cyrtoceras uncinatum Teijsm. & Binn. was described from plants cultivated in 
Bogor Botanic Garden originally collected from Sumatra by Teijsmann. No specimens 
have been found and it is possible that the description was based on a live plant. The 
name is neotypified with the lectotype of Hoya multiflora. 
In P herbarium there is only one specimen labeled in Costantin’s hand as Hoya 
reticulata Costantin, collected by Pierre on Mt. Krewanh. This specimen [P00700502] 
is therefore selected as lectotype. 
Centrostemma platypetalum Merr. was described based on Tso 23346. The NY 
duplicate is a fertile well-preserved specimen and is here selected as lectotype. 
HoyanabawanensisKloypQnb. &Wiberg,Fratema 15(2): 2 (2002).-TYPE: Originally 
from Malaysia, Sabah, Nabawan, cultivated in Sweden, Borlange, vouchered on 
unknown date (before March 2002), as Wiberg, E.K. s.n. (holotype UC [UC1776376]). 
Hoya nuttiana Rodda & Simonsson, Phytotaxa 140: 56 (2013). - TYPE: Malaysia, 
Sarawak, near Kuching, 24 October 1894, Haviland, G.D. & Hose , G.F. 8530 (holotype 
K [K000014447]). 
Hoya nyhuusiae Kloppenb., Fraterna 16(3): 1 (2003). - TYPE: Originally from 
Malaysia, Sabah, Mt. Kinabalu, cultivated in Sweden, Stockholm, vouchered in 2003 
as Nyhuus, T. s.n. (holotype UC [UC 1784974]). 
