Names and types of Bornean Hoya 
51 
(‘omlori- TYPE: Indonesia, Sumatra, Aceh, Gunung Leuser Reserve, Camp 
Simpang and vicinity, 19 August 1972, de Wilde, W.J.J.O & de Wilde-Duyfjes, B.E.E. 
14377 (holotype L; isotype K). 
Hoyapubera Blume, Bijdr. FI. Ned. Ind. 16: 1065 (1826). - Acanthostemmapuberum 
Blume, Rumphia 4: 29 (1849). - TYPE: [in m ari d m is insulae Nusae Kambangae ad 
arbores] (lectotype L [L0275692], designated here). 
Notes. A few specimens can be found in L labelled Hoya pubera Blume. Among these, 
[L0275692] bears a label, likely in Blume’s hand, with ‘1684 Dischidia elliptica Bl. 
Hoya pubera Bl.’ another specimen [L0275693] bears ‘ Acanthostemma pubera Bl, 
Hoya pubera Bl (illegible) Bl. 1684’, possibly in Blume’s hand. Both are sterile and 
cannot without doubt be linked to the protologue of Hoya pubera. Another specimen, 
in P [P00639851], was sent by Blume as Hoya pubera in 1836 and mentioned by 
Decaisne (1844: 639). Once again this is sterile and it is impossible to ascertain 
whether it was collected before the publication of H. pubera. Another specimen in 
U [U1102645] is labelled as collected by Blume, but is undated and just a fragment. 
Following Art. 9.12 of the ICN (McNeill et al., 2012), [L0275692] is to be considered 
as ‘remaining original material’ and is therefore here selected as the lectotype for Hoya 
pubera. 
Hoya phyllura O.Schwartz, Mitt. Inst. Bot. Hamburg 7: 26 (1931). - TYPE: 
Indonesia, Borneo, Lebang Hara, 24. November 1924, Winkler , H. 339 (lectotype 
HGB [HBG506353], designated here; isolectotype HBG [HBG506354]). 
Hoya linusii A.L.Lamb, Gavrus & Emoi, Sandakania 19: 50 (2014) [2 May 2014] . - 
TYPE: Originally from Malaysia, Sabah, Crocker Range, upper reaches of the Moyog 
River, cultivated at Kipandi Park, vouchered on 13 November 2007 as Gokusing, L in 
Lamb, A. 1223/2007 (holotype SAN; isotype SNP, n.v.). 
Notes. Schwartz cited only a single collection for Hoya phyllura , Winkler 339. Two 
duplicates are found in HBG. The first, bearing a sketch of the flowers is selected as 
the lectotype. 
Hoya ranauensis T.Green & Kloppenb., Hoya New 2(3): 21 (2014) [before 22 May 
2014]. -TYPE: Originally from Malaysia, Sabah, Ranau, Poring Hot Springs, cultivated 
in USA, Hawaii, Oahu, Ka‘a‘awa, garden of Ted Green, vouchered on unknown date 
as Green, T. s.n. (holotype BISH, sheet no. BISH757786 [BISH1049135]). 
Hoya retrorsa Gavrus, A.L.Lamb, Emoi & Gokusing, Sandakania 19: 59 (2014). - 
TYPE: Originally from Malaysia, Sabah, Pensiangan, Mount Lumatan, cultivated 
at Kipandi Park, vouchered on 4 May 2010 as Gokusing, L. in Lamb, A. 1950/2010 
(holotype SAN; isotype SNP n.v.). 
