Names and types of Bornean Hoya 
53 
Notes. The type of Hoya sipitangensis was cited as ‘52002 (UC), collected by Eva- 
Karin Wiberg et al. at Sipitang, Sarawak, Malaysia ex hort. Eva-Karin Wiberg at 
Borlange, Sweden’. The specimen labelled as holotype of Hoya sipitangensis in 
UC agrees in collecting information but does not bear the number 52002 which is, 
in any event, probably a collection date rather than a number as observed for Hoya 
walliniana and Hoya nyhuusiae. It is barcoded UC 1784973. As it is impossible to 
verily if [UC 1784973] is the intended holotype or another specimen obtained from the 
same plant in cultivation it is designated as neotype of Hoya sipitangensis. 
The collection date of the type of Hoya yapiana Kloppenb. is indicated in 
the protologue as ‘21 January 2008’ ; however, this is not the collection date of the 
specimen but the collection date of a living plant in Brunei that later bloomed in 
Singapore where it was cultivated by K.F. Yap, who made the type specimen from this 
plant on 20 April 2009. This collection date is mentioned on the holotype in SING but 
not on the duplicate in K. 
Hoya sigillatis subsp. sigillatis T.Green, Fraterna 17(3): 2 (2004). - TYPE: Originally 
from Malaysia, Sabah, Tenom Agricultural Park, cultivated in USA, Hawaii, 
Oahu, Ka‘a‘awa, vouchered on unknown date as Green, T. 91024 (holotype BISH 
[BISH1014783]). 
Hoya sigillatis T.Green subsp. paitanensis Gavras, A.L.Lamb, Emoi & Golcusing, 
Sandalcania 19: 68 (2014). - TYPE: Originally from Malaysia, Sabah, Beluran, 
Valley of the Paitan River, cultivated at Kipandi Park, vouchered on 10 April 2011 as 
Gokusing, L. & Lombika, J. in Lamb, A. 1757/2011 (holotype SAN; isotype SNPn.v.). 
Hoya spartioides (Benth.) Kloppenb., Fraterna 14(2): 8 (2001). - Astrostemma 
spartioides Benth., Hooker’s Icon. PI. 14: 7, t. 1311 (1880). - Absolmsia spartioides 
(Benth.) Kuntze, Revis. Gen. PI. 2: 418 (1891). - TYPE: Malaysia, North Borneo, 
1877-1878, Burbidge, F.W. s.n. (lectotype K [K000894739], designated by Wanntorp & 
Forster (2007), first step, designated here, second step; isolectotype K [K000894738]). 
Notes. Wanntorp & Forster (2007) selected Burbidge s.n. (K), a specimen not personally 
examined by the authors and mounted on two sheets, as lectotype of Hoya spartioides. 
However the two sheets are not clearly labelled as being part of the same specimen 
and must therefore be considered as two duplicates following ICN Art. 8.3 (McNeill 
et al., 2012). The specimen with barcode [K000894739] is therefore designated as the 
lectotype of Hoya spartioides. Syntype Motley , J. 978 is extant at K [K000894737]. 
Hoya telosmoides Omlor, Novon 6: 290 (1996). - TYPE: Malaysia, Sabah, Mt 
Kinabalu, Tenompok, 7 Jun 1932, Clemens, J. & M.S. 29828 (holotype BM; isotype 
K). 
Hoya undulata S.Rahayu & Rodda, Gard. Bull. Singapore 67: 86 (2015). - TYPE: 
Indonesia, West Kalimantan, Putussibau, 300-380 m, on a slope above a stream, 
February 2014, Sulaiman Hasim s.n. (holotype BO). 
