14 
Card. Bull. Singapore 69(1) 2017 
sometimes greenish maroon, shortly fused to the petiole, to 1.5 cm long, sparsely 
glandular hairy. Calyx green, clasping the corolla tube, lobes narrowly triangular or 
very narrowly ovate, 10-12 mm long, 8-10 mm wide, margin ciliate, outer surface 
glandular and eglandular hairy. Corolla very sparsely hairy outside and inside, tube 
usually cream or yellowish, sometimes whitish or pale purple, 2.4-5.5 cm long, 
slightly curved, 2-4 mm wide at the base, inside throat dark yellow to pale yellow, 
glandular hairs in the mouth of the tube above the anthers translucent pale yellow; 
upper lobes elliptic, not spreading, 4-5.5 x 5-6 mm, lower lateral lobes elliptic, 5-6 
x 4-6 mm, lower central lobe elliptic, slightly elongated, recurved or not, 6-8 x 5-10 
mm Stamens, filaments cream or white, inserted at 8-10 mm from the base of the 
corolla, 3-6.5 mm long, slightly curved or geniculate, glabrous; anthers pale yellow, 
pale brown or whitish, at base with long white deflexed hairs, anther thecae divergent, 
2-3 mm long; staminodes 2 (rarely 3), cream or green, vestigial, inserted at 9 mm 
from the base of the corolla tube, 2-7 mm long, glabrous. Nectary pale yellow or 
cream, a shallowly lobed ring, to 1 mm high. Pistil pale green; ovary 1.8-2.5 cm long, 
1 mm wide at the base, narrowing to 0.5 mm below the stigma, hairs glandular and 
eglandular; style 6-28 mm long; stigma deeply 2-lobed, to 3 mm long, above with 
very fine papillose hairs, beneath with very sparsely glandular hairs; ovules cream, 
less than 1 mm long. Capsules green, cylindric, slender, slightly curved towards tip, to 
8 cm long, 1.3-1.8 mm diameter, sparsely hairy; calyx persistent, green, hairy. Seeds 
yellowish cream to brownish, 435-550 x 100-230 pm, ellipsoid to narrowly ellipsoid, 
surface papillate or canaliculate. 
Distribution. Endemic to Malaysia: Peninsular Malaysia (Kedah, Pahang, Perak, 
Selangor and Terengganu) and Sarawak (Kuching Division). 
Habitat and ecology. Damp places on limestone karst hills, limestone cliffs, crevices 
and on open limestone rocks, on thin soil or on guano in caves or cave mouths, around 
the base or at the foot of limestone hills and at the base of shaded rocks. Altitude to 
240 m. 
ProvisionalIUCNconservation status. Least Concern (LC). Populations are widespread 
within the network of Totally Protected Areas in Taman Negara (the National Park), 
both in Pahang and Terengganu. 
Additional specimens examined. PENINSULAR MALAYSIA: Kedah: Kodiang, Bkt. Kaplu, 
6 Nov 2009, Rafidah FRI 64417 (KEP); ibidem, 27 May 2010, Rafidah FRI 64545 (KEP). 
Pahang: Bkt. Charas, 15 Oct 1931, Henderson 25233 (SING); ibidem, 26 Nov 1984, Kiew 
RK 1557 (KEP); ibidem, 1 Apr 2008, Rafidah FRI 55717 (KEP); Bkt. Chintamani, 4 Oct 1931, 
Henderson SFN 25033 (SING, BK); Gn. Jebak Puyuh, 10 Feb 1986, Kiew RK 2158 (KEP); 
ibidem, 10 Feb 1986, Kiew RK s.n. (KEP); Gn. Senyum, 30 Jul 1929, Henderson s.n. (SING); 
ibidem, 28 Nov 1984, Kiew RK 1587 (KEP); ibidem, 2 Apr 2008, Rafidah FRI 55721 (KEP); 
Gua Bama, 3 Apr 2008, Rafidah FRI 55726 (KEP); Gua Cermin R.F., 31 Mar 2008, Rafidah 
FRI 55713 (KEP); ibidem, 31 Mar 2008, Rafidah FRI 55714 (KEP); Gua Kechil, 30 Jul 2009, 
Rafidah FRI 643 79 (KEP); Kota Glanggi, 4 Aug 1929, Henderson SFN22419 (SING); ibidem, 2 
