Hoya of Papuasia I. 
123 
many other botanists (Vollering et al., 2016). The population size at the type locality 
was reduced from 2010 to 2012 due to expanding subsistence agriculture. 
Additional specimens examined. PAPUA NEW GUINEA: Morobe Province: Huon Peninsula, 
Hoboc, 1830 m, 24 December 2011, living accession NS 11-332, vouchered at Ukarumpa on 2 
June 2013 as Simonsson Juhonewe, N. & Juhonewe, F. NS0074L (LAE). 
Notes. Hoy a juhoneweana is easily distinguished from the large majority of Hoy a 
species due to the conspicuous sunken venation causing a characteristic bullate leaf 
surface, and by its dense cover of >1 mm long hairs on all vegetative parts, especially 
when young. Its leaves are, however, indistinguishable from Hoya urniflora. The two 
species can be separated when sterile because Hoya juhoneweana is pendulous while 
H. urniflora is a climber. When in flower, the two taxa are easily distinguished, as Hoya 
juhoneweana has >2.5 cm wide corollas whereas the corollas of H. urniflora are c. 1.5 
cm wide. Interestingly, the two species co-exist and intermingle at the same locality 
leading us to initially assume they were a single species during early expeditions. 
Upon flowering in cultivation, however, we realised there were two distinct species 
growing together in the same area. 
Hoya juhoneweana subsp. lindforsiana Simonsson & Rodda, subsp. nov. 
Similar to Hoya juhoneweana ssp. juhoneweana and Hoya urniflora in having sunken 
leaf venation and bullate leaf surface, but easily distinguished by the diameter of its 
corolla (4-5 cm), whereas it is 2.5-3 cm in H. juhoneweana subsp. juhoneweana and 
c. 1.5 cm in Hoya urniflora. - TYPE: Papua New Guinea, Madang Province, Mebu, 
disturbed primary riverine forest, 1300 m, living accession NS 12-279, vouchered 
at Ukarumpa on 9 March 2013, Simonsson Juhonewe, N. & Juhonewe, F. NS0068L 
(holotype LAE; isotype SING). (Fig. 12, 15) 
Slender pendulous epiphyte with occasional twining branches, with white latex in all 
vegetative parts. Stems up to 5 m long, flexible, often pendulous, cylindrical, 1-1.5 
mm in diameter, green, pubescent with very stiff, spreading, 0.5 mm or longer hairs; 
older stems lignified, up to 3 mm in diameter, greyish brown, pubescent or glabrous; 
intemodes 3-6(-25) cm long. Adventitious roots scattered along the stems. Leaves 
pendulous, petiolate; petiole terete, 2-10 x 1-1.5 mm, green, covered with very 
stiff, erect 1 mm or longer hairs; lamina elliptic-ovate, chartaceous, stiff and brittle 
when mature, 3-8 x 2-4 cm, bright green on adaxial surface, light green on abaxial 
surface, both surfaces covered by very stiff, erect, 1 mm or longer hairs, especially 
near base and abaxial midrib, apex cuspidate to acute, base round to attenuate, margin 
recurved; venation pinnate, midrib depressed on adaxial surface, secondary veins 
2-5 each side, evident and often depressed, giving the lamina a bullate appearance, 
anastomosing near the leaf margin, branching from the midrib at 45-80°; leaf colleters 
one pair at each lamina base, greyish white. Inflorescence one per node, extra-axillary, 
