Gardens’ Bulletin Singapore 69(1): 149-155. 2017 
doi: 10.3850/S2010098116000104 
149 
New combinations and typifications 
in Aglaomorpha (Polypodiaceae) 
S. Lindsay 12 , P.H. Hovenkamp 3 & D.J. Middleton 4 
‘Gardens by the Bay, 18 Marina Gardens Drive, Singapore 018953 
2 Present Address: Native Plant Centre, Horticulture and Community Gardening 
Division, National Parks Board, 1 Cluny Road, Singapore 259569 
stuart_lindsay @ nparks. go v. sg 
3 Naturalis Biodiversity Center, PO Box 2317, 2300 RA Leiden, The Netherlands 
4 Herbarium, Singapore Botanic Gardens, National Parks Board, 
1 Cluny Road, Singapore 259569 
ABSTRACT. In preparation for a new checklist of Singaporean plants, the Flora of Singapore 
project, and the Ferns of Thailand, Laos and Cambodia website, new combinations in 
Aglaomorpha Schott are made to accommodate species formerly placed in Drynaria (Bory) 
J.Sm. and Christiopteris Copel. Ten names are lectotypified, seven of which are second step 
lectotypifications, and one name is neotypified. 
Keywords. Christiopteris, Drynaria 
Introduction 
A close relationship between Aglaomorpha Schott and Drynaria (Bory) J.Sm. has long 
been suspected and led to the establishment of the tribe Drynarieae Subh.Chandra 
to accommodate these two genera (see Hennipman et al., 1990). On the other hand, 
the relationships of Christiopteris Copel. were presumed to be with other genera of 
Polypodiaceae (Hennipman et al., 1990) or were said to be uncertain (Hennipman et 
al., 1998). 
Recent phylogenetic research on these genera has shown that both Aglaomorpha 
and Christiopteris are nested within a paraphyletic Drynaria (Janssen & Schneider, 
2005; Schneider et al., 2008) which led Christenhusz & Schneider (2012) to propose 
the conservation of Drynaria against Aglaomorpha. After two rounds of voting 
no consensus could be reached as to whether to conserve or reject this proposal 
(Applequist, 2014). This means that the normal rules of priority should apply and 
that the name Aglaomorpha should be adopted for the genus sensu Into. Despite this, 
Christenhusz (2015) made combinations in Drynaria for all species in the clade which 
did not already have names in the genus. 
The Angiosperm Phytogeny Group has been deliberating on relationships and 
the classification of flowering plants since 1998 (APG, 1998; APG II, 2003; APG III, 
2009; APG IV, 2016) but only recently has there been an equivalent for lycophytes 
and ferns, the Pteridophyte Phylogeny Group (PPG I, 2016). This has set the standard 
for family delimitation in lycophytes and ferns based on a large synthesis of relevant 
