Hanguana podzolicola, a new record for Singapore 
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oblique, and stigma lobes which are connate at base and with sharp apices forming 
an equilateral triangle in H. triangulata). In addition the seed has a single, broadly 
and bluntly acute appendage (compared to the seed appendage bluntly bilobed in H. 
triangulata). Hanguana podzolicola is also similar in stature and inflorescence details 
to H. rubinea, but differs from it by its prominently corrugated leaves and abaxially 
more thickly flocculose indumentum (compared to the a lm ost flat lamina with a 
sparse indumentum in H. rubinea). They also differ in inflorescence branch length, 
fruit, and seed details (in Hanguana rubinea branches are up to 11 cm long, fruits are 
larger, 9-10 mm in diam., and turn ruby-red as they ripen, stigma is terminal or only 
slightly oblique, stigma lobes are similar to H. podzolicola , except larger, and the seed 
appendage is triangular). Hanguana podzolicola has the largest female inflorescences 
of any native Hanguana in Singapore. 
With the recent work on Hanguana in Singapore (Niissalo et al., 2014; Leong- 
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Skomickova & Boyce, 2015), the current paper brings the number of native Hanguana 
species in Singapore to five. Hanguana malayana (Jack) Merr., the name provisionally 
applied to the cultivated massive helophyte by Leong-Skornickova & Boyce (2015; 
see Fig. 1 & 11-12), is here updated to Hanguana anthelminthic a (Blume ex Schult. 
& Schult.f.) Masam. to reflect a recent clarification of the old names of Hanguana 
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(Leong-Skornickova & Niissalo, 2017). Hanguana malayana is a solitary forest 
species that is so far only known from Penang and has never been collected from 
Singapore, whereas Hanguana anthelminthica is the widespread massive helophyte 
that is also widely cultivated in the tropics, including in Singapore. An updated key to 
all Hanguana species in Singapore is provided. 
Hanguana podzolicola Siti Nurfazilah et al. - TYPE: Singapore, Mandai Road, 24 
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Aug 2015, Leong-Skornickova, J. HAN-76 (neotype SING [mounted over 2 sheets 
and including fruits preserved in spirit as part of a single specimen], here designated). 
(Fig. 1, 2) 
Herbaceous, dioecious mesophyte to c. 1.6 m tall; stem terete, to 3 cm in diam., 
basally semi-ascending, with age becoming leafless, terminally ascending with crown 
of up to 20 leaves; stolons absent. Leaves to 170 cm long, spreading then arching, 
bases imbricate margins hyaline (young leaves), turning erose-marcescent with age; 
pseudopetiole 60-70 cm long, c. 10-14 mm wide, accounting for 1/3—1/2 of entire 
leaf length, roundly channelled with sharp margins, strigose; leaf blade 80-105 x 
14-17 cm, narrowly elliptic, base attenuate, tip long, narrowly attenuate with apicule 
c. 4 mm, leathery, irregularly corrugated, adaxially mid to dark green, sparsely hairy 
(silky appressed hair; falling off in older leaves), abaxially lighter green when fresh, 
prominently flocculose (falling off in older leaves); midrib weakly impressed, of the 
same colour as the rest of the lamina adaxially, round-raised, lighter green, almost 
glabrous and shiny abaxially. Male inflorescences not observed, female inflorescences 
erect at anthesis. Female inflorescence/infructescence erect, comprising up to 9 
partial, whorled, alternate-secund, thyrsoid infructescences plus a terminal spike; 
