254 
Card. Bull. Singapore 69(2) 2017 
Caulescent herb to 60 cm tall, internodes 4-13 cm long, branches sometimes arising 
from the petiole of the basal cotyledon. Stem fleshy, pale green with purplish base, 
sparsely eglandular hairy. Leaves opposite, apart from the basal leaf; petioles pale 
green, 7-14 mm long; blades mid green above, paler beneath, ovate, 6.2-24 x 3.2-12.6 
cm, 1.7-2 times as long as wide, base cordate, apex acute to obtuse, eglandular hairy 
above and beneath, margin ciliate, margin entire to obscurely toothed, 8-15 pairs of 
secondary veins, venation sunken above and raised beneath in fresh material, flat in dry 
specimens, tertiary venation laxly reticulate. Inflorescences cristate, 10-16-flowered, 
all floral axes sparsely to densely eglandular hairy; peduncles 2-13 mm long, shortly 
fused with each other; bracts absent, except for first inflorescence arising from the 
petiole of each leaf, observed only in cultivated material; pedicels pale green to reddish 
brown, 3-26 mm long. Calyx actinomorphic, green, lobes succulent along the midrib, 
tube 1.3-3 mm, lobes imbricate, narrowly ovate, 7-18 x 2.9-3.6 mm, apex acute, 
margin entire, outside with eglandular hairs along the midrib, tip and margin, inside 
with sessile glands. Corolla 32-45 mm long, tube pale green at the base, then becoming 
dark orange, throat dark red with lateral purple spots, tube narrow at base, curved, 
then becoming broad, upper and lateral lobes spreading, tube glabrous at the base, 
then with mixed glandular (especially ventrally) and eglandular (especially dorsally) 
hairs, lobes papillose inside, the rest of the inside of the corolla glabrescent or sparsely 
glandular; tube 26-30 mm long dorsally, 29-32 mm ventrally, 22-27 mm between the 
lips; lobes with broadly obtuse apex and a minutely crenate margin, upper lobes 6-6.5 
x 10 mm, lateral lobes 8 x 12-13 mm, ventral lobe 10-11 x 12 mm. Stamens arising 
1.6-1.9 cm above the corolla base (the point where the corolla broadens), filaments 
slightly to strongly geniculate, 4.5-7 x 1 mm, pale yellow, glabrous, anthers white, 
hairy dorsally, 3-3.5 x 1.5-2.7 mm, apically joined by a connective, thecae divergent; 
lateral staminodes 0.5-2 mm long, arising 0.8-1.25 cm above the corolla base, central 
staminode 0.3-1 mm long, arising 0.9-1.35 cm above the corolla base. Disk annular, 
margin slightly lobed, 0.8-1 mm high. Pistil 27 mm long; ovary c. 1 cm long, 1.7 
mm diameter, glabrous in the bottom 1/3—1/2, densely hairy above; style slender, 1-2 
cm long, hairy; stigma bilobed, lobes c. 0.9-1 mm, papillose/plumose. Fruit green, 
4.5-6.2 cm long, 3.5^1 mm diameter, glabrous at the base, eglandular hairy apically, 
slightly curved. Seeds very dark brown, oval, 0.7-0.8 x 0.2-0.4 mm. 
Distribution. Central and Eastern Thailand. 
Habitat. On limestone. 
Provisional 1UCN conservation assessment. Vulnerable VU Blab(iii) + B2ab(iii). This 
species has an EOO and an AOO within the threshold to be considered endangered but 
is known from 5-10 localities, thereby making the Vulnerable status more appropriate. 
Additional specimens seen. Unknown province: 22 Oct. 1929, Kerr, A.F.G. (ABD); no label 
data (ABD); Cult. Hort. Bot. Reg. Kew., Nov 1926 “Bot. Mag. t. 9244” (K [2 sheets]); Cult. 
Hort. Bot. Reg. Kew. A.D. 196? (K); Cultivated, doubtfully from Mueang Ngao, Lampang, 
