156 
Two New Gregarines 
is carnivorous and lives upon Dasyhelea and several other dipterous larvae 
which are always found associated. 
The gregarine, in different stages of its development, occurs only in the 
midgut of its host, where it can remain for a long time as a trophozoite, 
attaining, at this stage, the size of a full-grown sporont, i.e. being 255 /x long 
and 18*5-20 /x wide. 
In all the stages the gregarine moves very slowly and easily bends and 
contracts its body (PI. X, figs. 11 and 12), which looks often as if it were 
composed of numerous rings; these curved specimens can frequently be seen 
performing a continual rotating movement. The epimerite of the cephalont 
-e 
A B 
Text-fig. 1. Dendrarhynchus system: A, the anterior portion of a trophozoite showing the epimerite (e). 
B, section of a trophozoite with its epimerite fixed in the epithelium of the host’s midgut. 
(Text-fig. 1, A and B) has the form of a disc surrounded by numerous more 
or less ramified papillae. 
At various stages, the cephalont, shedding off the epimerite, can separate 
itself from the host’s epithelial cell and become a free moving sporont (PI. X, 
figs. 8, 9 and 10). The body of the sporont is elongated with the posterior 
end slightly curved and of irregular contour; it does not seem to be divided 
into two segments, pro- and deutomerite, as is usual in cephaline gregarines. 
The ectoplasm is not very thick and shows well the longitudinally striated 
epicyte. The living, and especially the fixed and stained specimens, show, 
under the epicyte, a network of very well defined circular fibrils with oblique 
anastomoses which surround the whole body of the gregarine (Text-fig. 2). 
They undoubtedly correspond to the myocyte fibrils of Gregarina munieri 
described by Schneider (1875), although in this case the network is much 
