360 
Protozoa of Lizards , etc. 
granule there runs forward a fine rhizoplast. The deeply staining bodies which 
surround the nucleus vary considerably in number and arrangement. Some¬ 
times they are closely applied to the nuclear membrane while at others they 
are separated from it by a distinct interval. There may be many fine granules 
or a few larger masses. They may surround the nucleus or be limited to one 
side only or be entirely behind it. There may be only two bodies, one on each 
side of the nucleus, and of such a shape that they are swollen posteriorly and 
tapering anteriorly, with their anterior extremities nearer one another than 
the posterior ones. Prowazek (1904) notes these bodies and regards them as 
composed of chromatin material derived from the nucleus. I can find no 
evidence that these bodies arise from the nucleus. In those cases where they 
appear to be within the membrane careful observation shows that they are 
merely overlying it. Prowazek (1904) figures forms in which these bodies are 
on the inner surface of the membrane but in all the forms I have seen they are 
outside it even if lying against it. Prowazek’s idea that they are chromidial 
in nature was first contested by Dobell (1908), who wrote: “I think sufficient 
has been said to show that autogamy and chromidia are as yet unproven in 
the case-of Bodo.” They can best be spoken of as parabasal bodies similar 
to those which are associated with the origin of flagella in other flagellates. 
The rhizoplast arising from the blepharoplast or centrosome on the nuclear 
membrane passes forwards and is continued into the two flagella. At the 
extreme anterior end of the organism at the point where the flagella arise from 
the rhizoplast is occasionally seen a granule or thickening but I have been un¬ 
able to make out any structure in this (PL XX, Fig. 3, a , c, g). Prowazek (1904) 
figures the rhizoplast as terminating near the anterior end of the body in a 
sort of cone, beyond which is a second granule from which the flagella actually 
arise. If this structure is really present and there is a break in the rhizoplast 
then the flagellar origin is of a very specialized type. Is it not possible that 
this appearance is due to some peculiar plasticity of the anterior end of the 
body, which by retraction at the point of exit of the flagella gives rise to the 
cone-like appearance? At any rate in my preparations many of the flagellates 
seem to have a rhizoplast continued directly from the blepharoplast into the 
flagella. I have been unable to trace any connexion between the blepharo¬ 
plast and a granule within the karyosome of the nucleus such as’Prowazek 
describes. 
The anteriorly directed flagellum is often at least five times the length of 
the body of the elongate flagellates and in some of the smaller and ovoid 
forms it is even longer in proportion. The posteriorly directed flagellum 
reaches as much as three times the length of the body in the long forms. It 
is very much finer than the anteriorly directed flagellum and in some indi¬ 
viduals it appears to be attached to the body for a short distance (PI. XX, Fig. 
3, a , c, e), an attachment which may have to do with maintaining its direction. 
In many of the flagellates, however, there is no such attachment. 
