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THE WILSON ORNITHOLOGICAL SOCIETY 
FOUNDED 3 DECEMBER 1888 
Named after ALEXANDER WILSON, the first American ornithologist. 
President —Robert C. Beason, P. O. Box 737. Sandusky, OH 44871. USA; e-mail: Robert.C.Beason@ 
gmail.com 
First Vice-President Robert L. Curry, Department of Biology, Villanova University, 800 Lancaster 
Avenue. Villanova. PA 19085, USA; e-mail; robert.curry@villanova.edu 
Second Vice-President —Sara R. Morris, Department of Biology, Canisius College. 2001 Main Street, 
Buffalo. NY 14208, USA, e-mail: morris@canisius.edu 
Editorr-Clait E. Braun. 5572 North Ventana Vista Road. Tucson, AZ 85750, USA; e-mail: TWILSONJO@ 
comcast.net 
Secretary John A. Smallwood. Department of Biology and Molecular Biology. Montclair State University, 
Montclair. NJ 07043, USA; e-mail: smallwoodj@montclair.edu 
Treasurer —Melinda M. Clark, 526841 lighland Drive, South Bend IN 46635, USA; e-mail:wosclark@gmail.com 
Elected Council Members —Mary Bombcrger Brown, Mary C. Garvin. Timothy J. O'Connell, and Mark 
S. Woodrey (terms expire 2012); Mark E. Deutschlandcr and Rebecca J. Safran (terms expire 2013); 
William H. Barnard Margret I. Hatch, and Mark E. Hauber (terms expire2014). 
Living Past-Presidents- Pershing B. Hofslund, Douglas A. James, Jerome A. Jackson, Clait E. Braun, 
Richard C. Banks, Richard N. Conner, Keith L. Bildstein, Edward 11. Burtt Jr., John C. Kricher, William 
E. Davis Jr., Charles R. Blem. Doris .1. Watt, James I). Rising, and G. Dale Kennedy. 
Membership dues per calendar year are: Regular, $40.00; Student, $20.00; Family, $50.00; Sustaining, 
$100.00; Life memberships, $1,000.00 (payable in four installments). 
The Wilson Journal of Ornithology is sent to all members not in arrears for dues. 
THE WILSON JOURNAL OF ORNITHOLOGY 
(formerly The Wilson Bulletin) 
THE WILSON JOURNAL OF ORNITHOLOGY (ISSN 1559-4491) is published quarterly in March, June. 
September, and December by the Wilson Ornithological Society, 810 East 10th Street, Lawrence, KS 66044-8897. The 
subscription price, both in the United States and elsewhere, is $40.00 per year. Periodicals postage paid at Lawrence, KS. 
POSTMASTER: Send address changes lo OSNA. 5400 Bosque Boulevard. Suite 680, Waco, TX 76710. USA. 
All articles and communications for publication should be addressed to the Editor. Exchanges should be addressed 
to The Josselyn Van Tyne Memorial Library. Museum of Zoology. Ann Arbor. Ml 48109, USA. 
Subscriptions, changes of address, and claims for undelivered copies should he sent to OSNA. 5400 Bosque 
Boulevard. Suite 680, Waco, TX 76710, USA. Phone: (254) 399-9636; c-mail: busincssmosnabirds.org. Back issues or 
single copies are available for S12.00 each. Most back issues of the journal are available and may be ordered from OSNA. 
Special prices will be quoted for quantity orders. All issues of the journal published before 2000 are accessible on a free 
web site at the University of New Mexico library (http://clibrary.unm.edu/sora/). The site is fully searchable, and full-text 
reproductions of all papers (including illustrations) are available as cither PDF or DjVu files. 
€' Copyright 2012 by the Wilson Ornithological Society 
Printed by Allen Press Inc., Lawrence, KS 66044, USA. 
GOV FR. Wilson s Phalaropes (PhaUiropus tricolor). Illustration by Robin Corcoran. 
© This paper meets the 
requirements of ANSI/NISO Z39.48-1992 (Permanence of Paper). 
